Under this heading we describe the organs of circulation, by the action of which certain fluids are propelled through the body. It is customary to divide this branch of the subject into two sections, considering respectively the blood-vascular and lymphatic systems.
Blood-Vascular System.—This involves the consideration of the blood, a fluid which supplies nutriment to the tissues and receives effete material from them; the heart, a muscular organ which, by its contraction, initiates the motion of the blood; the arteries, a series of tubes which convey the blood from the heart to all parts of the body; the veins, tubes which return that fluid to the heart; and the capillaries, minute tubes joining the small arteries and veins.
Blood.—Blood is a fluid tissue, which nourishes all living structures, being the medium by which nutritive material is conveyed to, and effete or waste material conveyed away from the solid tissues. It is an opaque, thickish, clammy liquid, with a peculiar odor, sickly saline taste, and alkaline reaction. Its color varies in different parts of the same animal, that in the arteries being bright red or scarlet, while the blood in the veins is of a dark purplish hue.
When examined microscopically, the blood is found to consist of minute corpuscles, and a clear, transparent, yellow fluid, the liquor sanguinis, or plasma, in which the corpuscles float. The corpuscles are of two kinds, the red and the white or colorless; the former, by far more numerous, vary in proportion. Red corpuscles vary in shape, but in all mammals (animals that suckle their young) are more or less flat, the families excepted where they are oval, as in birds, reptiles, and fish, which are also nucleated. Their average diameter in the horse, ox or sheep is about 1/4000th part of an inch, their average thickness being about one-fourth of this. Each surface is depressed towards its center, hence the corpuscle is appropriately described as a bi-concaved disc.
The white corpuscles are larger than the red, round in shape, and nucleated.
The liquor sanguinis is pale and clear, and consists of water, fibrin, albumen, fatty compounds, extracts, odoriferous and saline matters. The serum is a thin, transparent liquid, of a pale-straw or yellow color, consisting of the liquor sanguinis deprived of fibrin. It contains nearly 90 per cent of water, is always slightly alkaline, and coagulates when heated, owing to the large quantity of albumen it contains. Fibrin is a white, stringy elastic substance, which, when the blood is in circulation, is in solution, and cannot be distinguished from the other constituents of the plasma.
Heart.—The heart is the principal organ of circulation; it weighs about six and one-half pounds in the average horse and acts as a force pump to force the blood through the arteries. It is composed of strong muscular tissue, which acts involuntarily, and is situated between the lungs, which are divided by what is known as the mediastinum. This is a division between the lungs made up of two folds, the heart being between them. The bottom or apex, of the heart is downward and rests just above the breast-bone; the upper part, or base is directed upward and to the left side, the left lung having a depression on its inner surface for the heart to work in. There is a covering or sack around the heart which helps to protect and support it in its place. It is attached above to the back-bone, and below to the bones of the breast. This sack is made up of fibrous tissue and is of a whitish appearance; inner surface is smooth, and supplied with numerous small glands which secrete an oily substance called serous fluid. This lubricates the outer surface of the heart and the inner surface of the sack so that in action it does not irritate the walls or surfaces. The cavity of the heart is divided into two parts, the right and left sides; each of these parts is again subdivided. The upper cavity is called auricle and the lower cavity ventricle; thus there are the right and left ventricle and right and left auricle. The right auricle communicates with the right ventricle by an opening in the septum or partition on the right side of the heart. This opening is guarded by a valve to keep the blood from flowing back into the auricle. The left auricle communicates with the left ventricle, same as on the right side. The right side of the heart is sometimes called the venous side and contains only venous or impure blood. The left side is sometimes called the arterial side. It contains pure blood only. This side of the heart is very much stronger and thicker than the right side.
THE HEART AND THE CHIEF BLOOD VESSELS OF THE HORSE—AFTER MEGNIN.