Scale and seal trees were abundant during the Pennsylvanian Period and were important contributors to coal beds. Although distantly related to the diminutive club-mosses and ground pines of the present, the trees grew on straight, slender trunks to heights of more than a hundred feet.
Reconstruction of Lepidodendron
(after Hirmer)
Scale trees were so called because their numerous, closely set, spirally arranged leaves left scarred “cushions” on the branches and trunk, making them appear scaly. Seal trees derived their name from the signetlike appearance of their leaf cushions. The two best known types belong to the genera Lepidodendron (scale tree) and Sigillaria (seal tree), and fossils of both are common in Illinois.
Lepidodendron had long, slender, somewhat tapering trunks. Some of the trees reached heights of more than 100 feet and measured more than two feet in basal diameter. The trunk ended in a spreading crown formed by repeated dichotomous branching. The leaves were awl-shaped or linear, ranging from one to 30 inches long.
The leaf cushions of Lepidodendron are diamond-shaped, longer than broad, and arranged in spiral rows around the trunk and branches. A different name, Lepidophyllum, is used for fossils of the long, bladelike leaf when it is found detached.
Spores were borne in long cylindrical cones at the tips of the branches. Those cones referred, or assigned, to the genus Lepidostrobus bore both small spores (microspores) and large spores (megaspores) in the same cones. Those in which only a large single spore, a somewhat seedlike structure, was developed in a spore sac (sporangium) are referred to the genus Lepidocarpon.
The rather commonly found genus Stigmaria comprises so-called “appendages” which, although stemlike in structure, apparently served as roots for the scale and seal trees. These appendages are identified by irregular spirals of circular scars (pits) that mark the attachment points of former rootlets.
Reconstruction of Sigillaria
(after Hirmer)