or,

1 ⁄ k = θ ⁄ log(r ⁄ a).

Then, if θ increase by 2π, while r increases to r1 ,

1 ⁄ k = (θ + 2π) ⁄ log(r1 ⁄ a),

which leads, by subtraction to

1 ⁄ k · log(r1 ⁄ r) = 2π.

Now, as α tends to 0, k (i.e. cot α) tends to ∞, and therefore, as k → ∞, log(r1 ⁄ r) → ∞ and also r1 ⁄ r → ∞.

Therefore if one whorl exists, the radius vector of the other is infinite; in other words, there is nowhere, even in the near neighbourhood of the pole, a complete revolution of the spire. Our spiral shells of small constant angle, such as Dentalium, may accordingly be considered to represent sufficiently well the true commencement of their respective spirals.

Let us return to the problem of how to ascertain, by direct measurement, the spiral angle of any particular shell. The method already employed is only applicable to complete spirals, that is to say to those in which the angle of the spiral is large, and furthermore it is inapplicable to portions, or broken fragments, of a shell. In the case of the broken fragment, it is plain that the determination of the angle is not merely of theoretic interest, but may be of great practical use to the conchologist as being the one and only way by which he may restore the outline of the missing portions. We have a considerable choice of methods, which have been summarised by, and are partly due to, a very careful student of the Cephalopoda, the late Rev. J. F. Blake[515]. {537}