Fig. 302.

Moreover, since the cavity below the septum is practically closed, and is filled either with air or with water, P will be constant over the whole area of the septum. And further, we must assume, at least to begin with, that the membrane constituting the incipient septum is homogeneous or isotropic.

Let us take first the case of a straight cone, of circular section, more or less like an Orthoceras; and let us suppose that the septum is attached to the shell in a plane perpendicular to its axis. The septum itself must then obviously be spherical. Moreover the extent of the spherical surface is constant, and easily determined. For obviously, in Fig. [302], the angle LCL′ equals the supplement of the angle (LOL′) of the cone; that is to say, the circle of contact subtends an angle at the centre of the spherical surface, which is constant, and which is equal to π − 2θ. The case is not excluded where, owing to an asymmetry of tensions, the septum meets the side walls of the cone at other than a right angle,

Fig. 303.

as in Fig. [303]; and here, while the septa still remain portions of spheres, the geometrical construction for the position of their centres is equally easy.

If, on the other hand, the attachment of the septum to the inner walls of the cone be in a plane oblique to the axis, then it is evident that the outline of the septum will be an ellipse, and its surface an {580} ellipsoid. If the attachment of the septum be not in one plane, but form a sinuous line of contact with the cone, then the septum will be a saddle-shaped surface, of great complexity and beauty. In all cases, provided only that the membrane be isotropic, the form assumed will be precisely that of a soap-bubble under similar conditions of attachment: that is to say, it will be (with the usual limitations or conditions) a surface of minimal area.

If our cone be no longer straight, but curved, then the septa will be symmetrically deformed in consequence. A beautiful and interesting case is afforded us by Nautilus itself. Here the outline of the septum, referred to a plane, is ap­prox­i­mate­ly bounded by two elliptic curves, similar and similarly situated, whose areas are to one another in a definite ratio, namely as

A1 ⁄ A2 = (r1r′1) ⁄ (r2r′2) = ε−4π cot α ,

and a similar ratio exists in Ammonites and all other close-whorled spirals, in which however we cannot always make the simple assumption of elliptical form. In a median section of Nautilus, we see each septum forming a tangent to the inner and to the outer wall, just as it did in a section of the straight Orthoceras; but the curvatures in the neighbourhood of these two points of contact are not identical, for they now vary inversely as the radii, drawn from the pole of the spiral shell. The contour of the septum in this median plane is a spiral curve identical with the original logarithmic spiral. Of this it is the “invert,” and the fact that the original curve and its invert are both identical is one of the most beautiful properties of the logarithmic spiral[535].