steps. And, since m and n are prime to one another (for otherwise the system would have been a composite or whorled one), it is evident that we can continue this process of convergence until we come down to a 1, 1 arrangement, that is to say to a leaf which is reached by a single step, in opposite directions from A and from a, which leaf is therefore the first leaf, next to A, of the fundamental or generating spiral. {646}

If our original lines along AO and aO contain, for instance, 13 and 8 steps respectively (i.e. m = 13, n = 8), then our next series, observable in the same cone, will be 8 and (13 − 8) or 5; the next 5 and (8 − 5) or 3; the next 3, 2; and the next 2, 1; leading to the ultimate condition of 1, 1. These are the very series which we have found to be common, or normal; and so far as our in­ves­ti­ga­tion has yet gone, it has proved to us that, if one of these exists, it entails, ipso facto, the presence of the rest.

In following down our series, according to the above construction, we have seen that at every step we have changed direction, the longer and the shorter sides of our triangle changing places every time. Let us stop for a moment, when we come to the 1, 2 series, or AT, aT of Fig. [326]. It is obvious that there is nothing to prevent us making a new 1, 3 series if we please, by continuing the generating spiral through three leaves, and connecting the leaf so reached directly with our initial one. But in the case represented in Fig. [326], it is obvious that these two series (A, 1, 2, 3, etc., and a, 3, 6, etc.) will be running in the same direction; i.e. they will both be right-handed, or both left-handed spirals. The simple meaning of this is that the third leaf of the generating spiral was distant from our initial leaf by more than the circumference of the cylindrical stem; in other words, that there were more than two, but less than three leaves in a single turn of the fundamental spiral.

Less than two there can obviously never be. When there are exactly two, we have the simplest of all possible arrangements, namely that in which the leaves are placed alternately on opposite sides of the stem. When there are more than two, but less than three, we have the elementary condition for the production of the series which we have been considering, namely 1, 2; 2, 3; 3, 5, etc. To put the latter part of this argument in more precise language, let us say that: If, in our descending series, we come to steps 1 and t, where t is determined by the condition that 1 and t + 1 would give spirals both right-handed, or both left-handed; it follows that there are less than t + 1 leaves in a single turn of the fundamental spiral. And, determined in this manner, it is found in the great majority of cases, in fir-cones and a host of other examples of phyllotaxis, that t = 2. In other words, in the {647} great majority of cases, we have what corresponds to an arrangement next in order of simplicity to the simplest case of all: next, that is to say, to the arrangement which consists of opposite and alternate leaves.

“These simple con­si­de­ra­tions,” as Tait says, “explain completely the so-called mysterious appearance of terms of the recurring series 1, 2, 3, 5, 8, 13, etc.[581] The other natural series, usually but misleadingly represented by convergents to an infinitely extended continuous fraction, are easily explained, as above, by taking t = 3, 4, 5, etc., etc.” Many examples of these latter series have been given by Dickson[582] and other writers.


We have now learned, among other elementary facts, that wherever any one system of helical spirals is present, certain others invariably and of necessity accompany it, and are definitely related to it. In any diagram, such as Fig. [326], in which we represent our leaf-arrangement by means of uniform and regularly interspaced dots, we can draw one series of spirals after another, and one as easily as another. But in our fir-cone, for instance, one particular series, or rather two conjugate series, are always conspicuous, while the others are sought and found with comparative difficulty.

The phenomenon is illustrated by Fig. [327], ad. The ground-plan of all these diagrams is identically the same. The generating spiral in each case represents a divergence of 3 ⁄ 8, or 135° of azimuth; and the points succeed one another at the same successional distances parallel to the axis. The rectangular outlines, which correspond to the exposed surface of the leaves or cone-scales, are of equal area, and of equal number. Nevertheless the appearances presented by these diagrams are very different; for in one the eye catches a 5 ⁄ 8 arrangement, in another a 3 ⁄ 5; and so on, down to an arrangement of 1 ⁄ 1. The math­e­mat­i­cal side of this very curious phenomenon I have not attempted to in­ves­ti­gate. But it is quite obvious that, in a system within {648} which various spirals are implicitly contained, the conspicuousness of one set or another does not depend upon angular divergence. It depends on the

Fig. 327.