In this sub-family there are numerous forms in which the elytra cover the pygidium, and in which the number of conspicuous ventral segments is reduced to five or even four. We use the term Coprides as equivalent to the "Laparosticti" of Lacordaire (Gen. Col. iii. 1856); it thus includes the "Coprini" and "Glaphyrini" of the Catalogus Coleopterorum, vol. iv. Munich, 1869.
Considérations genérales sur l'anatomic comparée des animaux articulés, etc., Paris 1828, 4to. xix. and 435 pp., and Atlas of ten (xx.) plates, and 36 pp.
Raspail, Mém. soc. zool. France, vi. 1893, pp. 202-213.
Ann. soc. ent. France, (v.) iv. 1874, p. 39.
In Theratides this outer lobe is in a rudimentary state, like a seta.