Fig. 440.—Starry Flounder, Platichthys stellatus (Pallas). Alaska.

The species of sole, about 150 in number, abound on sandy bottoms in the warm seas along the continents, very few being found about the Oceanic Islands. The three subfamilies, or tribes, may be designated as broad soles, true soles, and tongue-fishes.

The Broad Soles: Achirinæ.—The American soles (Achirinæ), or broad soles, resemble the smaller members of the turbot tribe of flounders, having the ventral fin of the eyed side extended along the ridge of the abdomen. The eyes and color are, however, on the right side. The eyes are separated by a narrow interorbital ridge. In most of these forms the body is broad and covered with rough scales. The species are mostly less than six inches long, and nearly all are confined to the warmer parts of America, many of them ascending the rivers. A very few (Aseraggodes, Pardachirus) are found in Japan and China. Some are scaleless and some have but a single small gill-opening on the blind side. The principal genus is Achirus. Achirus fasciatus, the common American sole, or hog-choker, is abundant from Boston to Galveston. Achirus lineatus and other species are found in the West Indies and on the west coast of Mexico. Almost all the species of Achirus are banded with black and the pectorals are very small or wanting altogether. All these species are practically useless as food from their very small size.

Fig. 441.—Hog-choker Sole, Achirus lineatus (L.). Potomac River.

The European Soles (Soleinæ).—The European soles are more elongate in form, with the ventral fins narrow and not extended along the ridge of the abdomen. The eyes are on the right side with no bony ridge between them. No species of this type is certainly known from American waters, although numerous in Europe and Asia. The species have much in common with the plaice tribe of flounders and may be derived from the same stock. One species, as above noted, is found in the Miocene.

The common sole of Europe, Solea solea, is one of the best of food-fishes, reaching a length, according to Dr. Gill, of twenty-six inches and a weight of nine pounds. As usually seen in the markets it rarely exceeds a pound. It is found from Norway to Italy, and when properly cooked is very tender and delicate, superior to any of the flounders. According to Dr. Francis Day, it appears to prefer sandy or gravelly shores, but is rather uncertain in its migrations, for, although mostly appearing at certain spots almost at a given time, and usually decreasing in numbers by degrees, in other seasons they disappear at once, as suddenly as they arrive. Along the British seacoast they retire to the deep as frosts set in, revisiting the shallows about May if the weather is warm, their migrations being influenced by temperature. The food of the sole is to a considerable extent molluscous, but it is also said to eat the eggs and fry of other fishes and sea-urchins.

The spawning season is late in the year and during the spring months. The ova are in moderate number; a sole of one pound weight has, according to Buckland, about 134,000 eggs. The newly hatched, according to Dr. Day, do not appear to be commonly found so far out at sea as some other species. They enter into shallow water at the edge of the tide and are very numerous in favorable localities.

As is well known, the sole is one of the most esteemed of European fishes. In the words of Dr. Day, "the flesh of this fish is white, firm, and of excellent flavor, those from the deepest waters being generally preferred. Those on the west coast and to the south are larger, as a rule, than those towards the north of the British islands. In addition to its use as food, it is available for another purpose. The skin is used for fining coffee, being a good substitute for isinglass, and also as a material for artificial baits.