"(8) The absence of a pecten, and presence of a vascular membrane between the vitreous humor and the retina.
"(9) The opisthocœlous form of the vertebræ.
"(10) The articulation of the ventral parts of the hæmal arches of the tail with the processes of the vertebral column.
"(11) The absence of a division of the muscles into dorso-lateral and ventro-lateral divisions.
"(12) The complete segmentation of the ovum.
"The list just given appears to us sufficient to demonstrate that Lepidosteus cannot be classed with the Teleostei; and we hold that Müller's view is correct, according to which Lepidosteus is a true Ganoid.
"The existence of the Ganoids as a distinct group has, however, recently been challenged by so distinguished an ichthyologist as Günther, and it may therefore be well to consider how far the group as defined by Müller is a natural one for living forms, and how far recent researches enable us to improve upon Müller's definitions. In his classical memoir the characters of the Ganoids are thus shortly stated:
"'These fishes are either provided with plate-like angular or rounded cement-covered scales, or they bear osseous plates, or are quite naked. The fins are often, but not always, beset with a double or single row of spinous plates or splints. The caudal fin embraces occasionally in its upper lobe the end of the vertebral column, which may be prolonged to the end of the upper lobe. Their double nasal openings resemble those of Teleostei. The gills are free, and lie in a branchial cavity under an operculum, like those of Teleostei. Many of them have an accessory organ of respiration, in the form of an opercular gill, which is distinct from the pseudobranch, and can be present together with the latter; many also have spiracles like Elasmobranchii. They have many valves in the stem of the aorta like the latter, also a muscular coat in the stem of the aorta. Their ova are transported from the abdominal cavity by oviducts. Their optic nerves do not cross each other. The intestine is often provided with a spiral valve, like Elasmobranchii. They have a swimming-bladder with a duct, like many Teleostei. Their pelvic fins are abdominal.
"'If we include in a definition only those characters which are invariable, the Ganoids may be shortly defined as being those fish with numerous valves to the stem of the aorta, which is also provided with a muscular coat, with free gills, and an operculum, and with abdominal pelvic fins.'
"To these distinctive characters he adds, in an appendix to his paper, the presence of the spiral valve, and the absence of a processus falciformis and a choroid gland.