The ordinary Mendelian pictures a unit character in a cross that obeys Mendel’s law, as follows:—
| D |
| R, |
| D(R) |
| R(D), |
| R(D) |
| R(D). |
| R(R) |
| R(R). |
Unit Characters
Mendelian phenomena force upon us the conclusion that organisms display a number of unit characters, each of which behaves in much the same way as a radicle does in chemistry, inasmuch as for one or more of these characters others can be substituted without interfering with the remaining unit characters. For example, it is possible to replace the chemical radicle NH3 by the radicle Na2; e.g. (NH3)2SO4 (ammonium sulphate) may be transformed into Na2SO4 (sodium sulphate).
The conclusion that each organism is composed of a number of unit characters, which sometimes behave more or less independently of one another, is one which most biologists who have studied the phenomena of inheritance appear to have arrived at. Zoologists are mostly of opinion that these characters, or rather their precursors, exist as units in the fertilised egg. Very varied have been the conceptions of the nature of these biological units. Almost every biologist has given a name to his particular conception of them. Thus we have the gemmules of Darwin, the unit characters of Spencer, the biophors of Weismann, the micellæ of Naegeli, the plastidules of Haeckel, the plasomes of Wiesner, the idioblasts of Hertwig, the pangens of De Vries, and so on. It is unnecessary to extend this list. It must suffice that almost every investigator of the phenomena of inheritance believes in these units, and calls them by a different name. Moreover, each clothes them with characteristics according to his taste or the fertility of his imagination.
Chemical Molecules
These units behave in such a way as to suggest to us an analogy between them and the chemical molecules. The sexual act would appear to resemble a chemical synthesis in some respects. One of the most remarkable phenomena of chemistry is that of isomerism. It not infrequently happens that two very dissimilar substances are found, upon analysis, to have the same chemical composition, that is to say, their molecules are found to be composed of the same kind of atoms and the same number of these. Thus chemists are compelled to believe that the properties of a molecule are dependent, not only on the nature of the atoms which compose it, but also on the arrangement of these within the molecule. To take a concrete example: Analysis shows that both alcohol and ether are represented by the chemical formula C2H6O. In other words, the molecule of each of these compounds is made up of two atoms of the element Carbon, six of the element Hydrogen, and one of the element Oxygen. Now, every chemical atom possesses the property which chemists term valency, in other words, the number of other atoms with which it can directly unite is strictly limited. All atoms of the same element have the same valency. Monovalent atoms are those which can, under no circumstances, unite with more than one other atom. The Hydrogen atom is an example of such an atom. Divalent atoms, as, for example, that of Oxygen, can unite with one other atom of similar valency or with two monovalent atoms. Similarly, a trivalent atom, such as that of Nitrogen, can unite with three monovalent atoms. A tetravalent atom, such as that of Carbon, can combine with four monovalent atoms. There are also pentavalent and hexavalent atoms. Now, by indicating the valency of any given atom by a stroke for each monovalent atom with which it is able to combine, chemists have been able to represent the molecule of every compound, or, at any rate, of every inorganic compound, by what is known as a graphic or structural formula. Thus, ethylic alcohol is represented by the formula:—