In the early part (winter and spring) of 1939, at Ames, Iowa, Polderboer, Kuhn and Hendrickson (1941) studied four weasels living in four separate dens on 160 acres typical of Iowa farmland and excavated three of the dens. One den was in a weed patch in an old mole run. The nest chamber, approximately nine inches in diameter and six inches below the surface of the ground "was filled with grasses packed in a layer-like formation. In the center of this mass was a nest hollow lined with patches of mouse and shrew fur. Beneath this layer of fur and at the sides of the nest were skins, various bones, and skulls of partially eaten mice and shrews . . . scats [were in the nest]. . . . At intervals, layers of clean grass had been laid over the filth of the former bed, thus giving the nest a stratified appearance." A second den, of a large male, was in a field of sweet clover two feet high in the former burrow of a Franklin's ground squirrel. The nest cell, seven inches in diameter and nine inches below the surface of the ground, "was lined with grasses mixed with much rabbit and mouse fur. Some scats, and bones and fur of mice and shrews were matted together in layers at the bottom of the nest." When this den was abandoned the male weasel occupied, for a month, another burrow, 20 rods distant, of a Franklin ground squirrel, in the field of sweet clover. The nest cell measured 11 by nine inches and was 11 inches below the surface of the ground. "Two nest layers were present. The first, composed chiefly of coarse straw and grass, had apparently been occupied at some time by a spotted skunk. . . . On top of the skunk nest was the weasel nest composed of fine grasses, mouse fur, and skeletal remains of mice."

Relation of the Sexes to each other and to the young

Quick (1944:75) writes that on March 28, in Michigan, he found the tracks of a male and those of a smaller animal, supposedly a female, meeting. The two "then led along the fence for about 18 chains and both entered the den of the male. . . . Only the tracks of the smaller weasel left the den on the same date. Observation on April 12 showed that the large male still occupied the den." I am at a loss to explain this behavior since breeding would not be expected to occur in late March and since I suppose that the male and female do not live together except in the breeding season. Consequently, I wonder if the sign was wrongly read.

[Fig. 29.] Map showing the geographic ranges of the subspecies of Mustela frenata and Mustela africana.

1. M. f. noveboracensis
2. M. f. occisor
3. M. f. primulina
4. M. f. arthuri
5. M. f. olivacea
6. M. f. peninsulae
7. M. f. spadix
8. M. f. longicauda
9. M. f. oribasus
10. M. f. alleni
11. M. f. arizonensis
12. M. f. nevadensis
13. M. f. effera
14. M. f. washingtoni
15. M. f. saturata
16. M. f. altifrontalis
17. M. f. oregonensis
18. M. f. munda
19. M. f. xanthogenys
20. M. f. nigriauris
21. M. f. latirostra
22. M. f. pulchra
23. M. f. inyoensis
24. M. f. neomexicana
25. M. f. texensis
26. M. f. frenata
27. M. f. leucoparia
28. M. f. perotae
29. M. f. macrophonius
30. M. f. goldmani
31. M. f. tropicalis
32. M. f. perda
33. M. f. nicaraguae
34. M. f. costaricensis
35. M. f. panamensis
36. M. f. meridana
37. M. f. affinis
38. M. f. aureoventris
39. M. f. helleri
40. M. f. macrura
41. M. f. agilis
42. M. f. boliviensis
43. M. a. africana
44. M. a. stolzmanni

Hamilton (1933:328), however, writes that M. f. noveboracensis is to "be found in pairs when caring for the young. During mid-May, 1927, I several times saw a male of this species carrying food to a den of young ones." Green (1936), in May in Gratiot County, Michigan, remarks that while he was uncovering and examining a nest of four young weasels, two adults ran about excitedly and one removed a young weasel. In instances where several nearly full-grown young have been obtained from one den it has been my experience (Hall, 1946:191) that the only adult trapped there was the female; no adult male was found or in the one instance when found he was living alone in a den 200 yards away from the den of the female and her young. Data are too few to warrant a definite conclusion about the extent to which the male aids in rearing the young, but I have wondered if he might not do so when the young were less than half grown and then live alone when they were more than half grown.

Mustela frenata noveboracensis (Emmons)