A subadult female, of abnormal color, no. 47149, Mus. Vert. Zoöl., taken by Mr. H. E. Wilder at Carlotta, California, on December 20, 1930, in a region where weasels do not turn white in winter, is white, except for the black tip of the tail, but has a suffusion of orange. This specimen, discussed at greater length on page [43], is instructive in that it suggests that there are separate determiners for the brown and red elements of the pelage. It is interesting also as suggesting how natural selection may tend to eliminate from the population a conspicuous color-variation of this kind. At any rate, Mr. Wilder (Ms.) states: "This specimen was picked up in a field, where it evidently had been dropped by a hawk or an owl." The braincase of the skull is crushed in three places as though by a raptor's beak. None of the several other weasels, all normally colored, saved by Mr. Wilder from this general locality gives evidence of having fallen a victim to a raptor.
Only 2 skulls of the 12 adults and subadults examined show malformation of the frontal sinuses such as results from the presence of parasites.
Specimens examined.—Total number, 29, arranged within states from north to south by counties. Unless otherwise indicated specimens are in the collection of the United States National Museum.
California. Del Norte County: Requa, 1[8]. Humholdt County: Eureka, 2 (1[74], 1[75]); Ferndale, 1[74]; Fortuna, 2[63]; Carlotta, 6 (3[74], 3[59]); 12 mi. E Bridgeville, 1[59]; 2 mi. W Bridgeville, 1[59].
Oregon. Washington County: Forest Grove, 1. Marion County: Salem, 2. Benton County: Buchanan, 1. Lane County: McKenzie Bridge, 1[101]; Vida Fish Hatchery, 1[101]; Eugene, 1[101]. Douglas County: Anchor, 1. Curry County: Gold Beach, 2[60]. Josephine County: Rogue River Valley (Grants Pass), 1; 13 mi. SW Grants Pass, 1[101]. Jackson County: Medford, 2; 6 mi. S Medford, 1[74].
Mustela frenata munda (Bangs)
Long-tailed Weasel
Plates [1], [19], [20], [21], [22], [23], [30], [34], [35], [36] and [40]
Putorius xanthogenys mundus Bangs, Proc. New England Zoöl. Club, 1:56, June 9, 1899; Stephens, California mammals, p. 247, 1906.
Mustela frenata, Audubon and Bachman, Journ. Acad. Nat. Sci. Philadelphia, 8 (Pt. 2):291, 1842 (North California about 40° latitude).
Mustela xanthogenys munda, Miller, U. S. Nat. Mus. Bull., 79:99, December 31, 1912.
Mustela frenata munda, Hall, Carnegie Instit. Washington Publ. 473:107, November 20, 1936.
Type.—Male, adult, skull, os penis and skin; no. 5459, collection of E. A. and O. Bangs, but now in collection of Mus. Comp. Zoöl.; Point Reyes, Marin County, California; June 19, 1896; obtained by C. A. Allen; original no. 931. (See comments under "Remarks," below, on places in California to which the name Point Reyes has been applied.)
The skull (pls. 19-21, 30) is complete and unbroken. I1 on each side and right I2 are broken away; p2 and p3 on each side have been aborted and the only alveoli remaining are two for the right p3. Otherwise all teeth are present and entire. The skin is fairly well made and in good condition.
Cranially, the type is a "runt"; its small size and the circumstance that the tympanic bulla is longer than the lower molar and premolar tooth-row and longer than the rostrum are features which differentiate the type from any other specimen seen of this race.
Range.—Sea level to at least 6,000 feet (South Yolla Bolly Mountain, Trinity County, California); Upper Sonoran and Transition life-zones of the coast and Coast Range of northwestern California from the Golden Gate northward into southern Humboldt and Trinity counties. See figures [29] and [30] on pages 221 and 314.
Characters for ready recognition.—Differs from M. f. oregonensis in longer hind foot of males which is more than 50 mm., and in both sexes, larger, more prominently ridged skull (see measurements and plates); from M. f. saturata by presence of nasofrontal white spot, larger and relatively shallower skull of males and larger skull of female; from M. f. nevadensis by presence of well-developed, white, facial markings; absence of color of underparts on ventral face of proximal third of tail; and hind foot of males more than 50; from M. f. xanthogenys by near (l) Sudan Brown to near (l) Antique Brown rather than Buckthorn Brown colors of upper parts and greater size, and in adult male basilar length more than 45 and hind foot more than 47; from M. f. nigriauris by having inside of ears same color as back rather than much darker than back.
Description.—Size.—Male: Three adults and two young from Point Arena and Gualala, Mendocino County, yield average and extreme measurements as follows: Total length, 447 (434-470); length of tail, 167 (150-185); length of hind foot, 53 (50-60). Corresponding measurements of three adults from 5 and 6 miles west of Inverness, Marin County, are: 430 (420-440), 154 (141-160), 48 (48-49). Corresponding measurements of four individuals (3 adults and 1 young of large size) from South Yolla Bolly Mountain, Trinity County, are: 383 (374-400), 134 (130-138); 44 (43-44). The tail averages 60 per cent as long as the head and body in the series from Point Arena, 56 per cent in the series from Point Reyes, and 53 per cent in the series from South Yolla Bolly Mountain. In every specimen except two, length of hind foot less than basal length. The two exceptions are no. 19720, M.V.Z., male adult from Point Arena in which the hind foot is recorded as 60 (probably an error in measurement), and no. 19721, M.V.Z., from the same place, in which the skull has not yet attained its full growth.
Female: One adult from Point Arena measures as follows: Total length, 383; length of tail, 134; length of hind foot, 43. Corresponding measurements of an adult from seven miles north of Laytonville, Mendocino County, are: 336, 121, 33 (= 36 on dried skin). Corresponding measurements of an adult from South Yolla Bolly Mountain, Trinity County, are, 326, 113, 37. In these three specimens, the tail is, in the order given, 54, 56, and 53 per cent as long as the head and body. Length of hind foot more than basal length.
Differences in external measurements of the two sexes as indicated by the five males and one female from Point Arena, are: Total length, 64; length of tail, 33; length of hind foot, 10. Weights of 2 adult males are 265 and 221 grams and of one adult female 155 grams.
Externals.—As described in Mustela frenata nigriauris.
Color.—Spot between eyes, narrow band or spot confluent with color of underparts on each side of head anterior to each ear, chin, lower lips, and rarely posterior third or less of each upper lip white; dark spot posterior to each angle of mouth uniformly present and of large size; tip of tail black; remainder of upper parts near (14 l) Sudan Brown and tone 4 of Raw Umber of Oberthür and Dauthenay, pl. 301; occasionally, slightly darker brown on forehead, nose, and about eyes. Underparts near (a to c) Ochraceous-Buff and sometimes Orange-Buff. Color of underparts extends distally on posterior sides of forelegs over toes onto antipalmar faces of feet and wrists, on medial sides of hind limbs over antiplantar faces of toes. Least width of color of underparts averaging, in a series of 5 males from Mendocino County, 57 (46-67) per cent of greatest width of color of upper parts; 38 (35-40) in 3 males from Point Reyes, Marin County. Black tip of tail in Mendocino County series averaging 53 (46-60) mm., which is same length as hind foot and 32 per cent of length of tail. In Point Reyes males, black tip of tail averages 44 (34-52) mm., which is less than length of hind foot and 45 per cent as long as tail-vertebrae.
Several specimens of the smaller, inland variant (see under "Remarks") are near (l) Antique Brown rather than near (14 l) Sudan Brown above and hence do not differ in this respect from nigriauris.
Skull and teeth.—Male (based on 3 adults from Mendocino County): See measurements and plates [19]-[23], [30]. As described in Mustela frenata nigriauris except that: Weight, 6.0 (5.4-6.3) grams; basilar length, 47.6 (46.5-48.2); length of tympanic bulla more than length of lower molar and premolar tooth-row.
Female (based on no. 19723, M.V.Z., from Point Arena): See measurements and plates [34]-[36], [40]. As described in M. f. nigriauris except that: Weight, 3.0 grams; basilar length, 42.3.
The skull of the female is 50 per cent lighter than that of the average male.