[13] E. J. Marey, ‘Analyses of the Movements of the Horse by the Chromophotograph’ (La Nature, June 11, 1898).

Fig. 42.—Extension of the Humerus: Right Anterior Limb of the Horse, External Surface (after a Chromophotographic Study by Professor Marey).

The Articulation of the Elbow, or the Humero-ulnar Articulation.—In this articulation, which is constructed in the form of a true hinge, the movements of flexion and extension alone are possible. In flexion, the forearm, directed forwards, is folded on the arm, with which, in certain circumstances, it comes in contact. For example, in a horse of mettle which leaps over an elevated obstacle, the animal forcibly raises his fore-limbs by flexing them. Flexion is produced to the same extent, and even more so, and for a longer period, in felides which crouch.

In extension, on the contrary, the forearm is carried backward. This movement being limited only by the contact of the tip of the olecranon with the bottom of the olecranon fossa of the humerus, the forearm is enabled, in this case, to move until it is in line with the arm. For example, during walking, when one of the anterior limbs, having reached the end of its resting stage, is considerably inclined downwards and backwards.

The apex of the olecranon process—that is to say, the point of the elbow—forms a marked prominence, more salient in flexion than in extension, as in the corresponding region of the human elbow.

The Radio-ulnar Articulation.—It is in the dog and the cat, in which the two bones of the forearm articulate by their extremities only, and remain separate in the rest of their extent, that the articulations call for special notice.

In the upper part, the radius rotates on itself; while below, it rotates around the ulna. It follows that the forearm, which in all quadrupeds is in a state of permanent pronation, can, in carnivora, take the position of supination, or rather, of demi-supination. In fact, whatever be the mobility of the two bones of the forearm, the movement is not able to bring the palmar surface to the front, but only to direct it towards the median line.

The Articulation of the Wrist.—Here are found, as in man, three superimposed articulations: the radio-carpal, intercarpal, and carpo-metacarpal.