a Chorion. b The larger absorbent extremities, the site of the placenta. c Allantois. d Amnion. e Urachus. é Bladder. f Vesicula umbilicalis. g Communicating canal between the vesicula umbilicalis and intestine. h Vena umbilicalis. i i Arteriæ umbilicales. l Vena omphalo-meseraica. k Arteria omphalo-meseraica. n Heart. o Rudiment of superior extremity. p Rudiment of lower extremity. From Carus.

The hands seem to be fixed to the shoulders without arms, and the feet to adhere to the ossa illi; the liver seems to fill the whole abdomen; the ossa innominata, the ribs, and scapulæ are cartilaginous.

In a short time the little stump-like prominences of the extremities become longer, and are now divided into two parts, the superior into the hand and the fore arm, the inferior into the foot and leg; in one or two weeks later, the arms and thighs are visible. These parts of the extremities which are formed later than the others, are at first smaller, but as they are gradually developed they become larger. When the limbs begin to separate into an upper and lower part, their extremities become rounder and broader, and divided into the fingers and toes, which at first are disproportionately thick, and until the end of the third month are connected by a membranous substance analogous to the webbed feet of water birds; this membrane gradually disappears, beginning at the extremities of the fingers and toes, and continuing the division up to their insertion. The external parts of generation, the nose, ears, and mouth appear after the development of the extremities. The insertion of the umbilical cord changes its situation to a certain degree; instead of being nearly at the inferior extremity of the fœtus as at first, it is now situated higher up on the anterior surface of the abdomen. The comparative distance between the umbilicus and pubis continues to increase, not only to the full period of gestation, when it occupies the middle point of the length of the child’s body, as pointed out by Chaussier, but even to the age of puberty, from the relative size of the liver becoming smaller.

Though the head appears large at first, and for a long time continues so, yet its contents are tardy in their development, and until the sixth month the parietes of the skull are in great measure membranous or cartilaginous. Ossification commences in the base of the cranium, and the bones under the scalp are those in which this process is last completed.

The contents of the scull are at first gelatinous, and no distinct traces of the natural structure of the brain can be identified until the close of the second month; even then it requires to have been sometimes previously immersed in alcohol to harden its texture. There are many parts of it not properly developed until the seventh month. In the medulla spinalis no fibres can be distinguished until the fourth month. The thalami nervorum opticorum, the corpora striata, and tubercula quadrigemina, are seen in the second month; in the third, the lateral and longitudinal sinuses can be traced, and contain blood. In the fifth we can distinguish the corpus callosum; but the cerebral mass has yet acquired very little solidity, for until the sixth month it is almost semi-fluid. (Campbell’s System of Midwifery.)

About the end of the third, during the fourth, and the beginning of the fifth months, the mother begins to be sensible of the movements of the fœtus. These motions are felt sooner or later, according to the bulk of the child, the size and shape of the pelvis, and the quantity of fluid contained in the amnion, the waters being in larger proportionate quantity the younger the fœtus.

The secretion of bile, like that of the fat, seems to begin towards the middle of pregnancy, and tinges the meconium, a mucous secretion of the intestinal tube which had hitherto been colourless, of a yellow colour. Shortly after this the hair begins to grow, and the nails are formed about the sixth or seventh month. A very delicate membrane (membrana pupillaris,) by which the pupil has been hitherto closed, now ruptures, and the pupil becomes visible. The kidneys, which at first were composed of numerous glandular lobules (seventeen or eighteen in number,) now unite, and form a separate viscus on each side of the spine; sometimes they unite into one large mass, an intermediate portion extending across the spine, forming the horse-shoe kidney.

Lastly, the testes, which at first were placed on each of the lumbar vertebræ, near the origin of the spermatic vessels, now descend along the iliac vessels towards the inguinal rings, directed by a cellular cord, which Hunter has called Gubernaculum testis: they then pass through the openings carrying before them that portion of the peritoneum which is to form their tunica vaginalis.

The length of a full-grown fœtus is generally about eighteen or nineteen inches; its weight between six and seven pounds. The different parts are well developed and rounded; the body is generally covered with the vernix caseosa;[28] the nails are horny, and project beyond the tips of the fingers, which is not the case with the toes; the head has attained its proper size and hardness; the ears have the firmness of cartilage; the scrotum is rugous, not peculiarly red, and usually containing the testes. In female children the nymphæ are generally covered entirely by the labia, the breasts project, and in both sexes frequently contain a milky fluid. As soon as a child is born, which has been carried the full time, it usually cries loudly, opens its eyes, and moves its arms and legs briskly; it soon passes urine and fæces, and greedily takes the nipple. (Naegelé’s Hebammenbuch.)

Thus, then, in the space of forty weeks, or ten lunar months, from an inappreciable point, the fœtus attains a medium length of about eighteen or nineteen inches, and a medium weight of between six and seven pounds. As these observations on the development of the ovum show that the structural arrangement of the embryo undergoes a succession of changes, by which it gradually rises from the lowest to the highest scale of formation, so we shall find it furnished with a succession of means for its nutrition, each corresponding more or less to the particular grade of development which it may have attained. Its earliest source of nourishment is doubtless the vitellus, or albuminous contents of the vesicula umbilicalis. The radicle or primitive trace, in this respect, bears a strong analogy to the seed of a plant; it brings with it its own supply of nourishment for its first stage of growth; in the latter, the cotyledons afford nourishment to the little plumula, until, by the formation of roots and absorption of moisture from the surrounding soil, it is enabled to support the early rudiment of the future plant. The early function of the chorion is very analogous to that of roots; it is an absorbing apparatus, collecting nourishment by means of its numerous absorbing fibrillæ: hence, according to Lobstein, the umbilical vein exists for some time previous to the umbilical arteries, and seems to perform an office in the fœtus similar to that of the thoracic duct at a later period; its radicles or absorbing extremities seem to absorb a milky fluid, which after the first two months is found in the placenta, and which must be looked upon as a means of nourishment which does not exist in the latter months. This milky fluid was noticed by Leroux, who even then expressed his doubts, whether the radicles of the umbilical vein receive blood from the mother, or whether they only serve to absorb a white fluid which resembles chyle. In some manuscript notes of Dr. Young’s lectures, which were taken by the late Dr. Parry, of Bath, when a student at Edinburgh, we find the following observation: “There is evidently in the placenta, besides blood-vessels, some other substance, which serves to absorb juices from the uterus, and to convert these into a chylous matter proper to nourish the fœtus, and this matter is absorbed by the umbilical veins. This seems to be proved from the consideration of the placenta of animals which have cotyledons; for, on squeezing these glandular substances, we force out a sort of chylous liquor, and these are surrounded by the placenta, which absorb their liquor and convey it to the fœtus.”