Cuvier divided the Linnean genus into two subgenera, Mustela and Putorius, the first, Martens and Sables, possessing an additional small premolar on each side of the jaw; the second including the Polecats, Stoats, and Weasels. At a later date Nilsson called these subgenera genera, substituting the name Martes for the Martens and giving that of Mustela to the Weasels. This has the effect of making the name of the Pine Marten, Martes martes, which is rather ridiculous; and we have preferred to retain the Linnean name Mustela martes. The old spelling of the popular name was Martin, but in recent works, to avoid any possible confusion with the birds of that name, zoologists have agreed to use e as the second vowel when writing of the mammals.
The Pine Marten is found in all the wooded regions of Europe and into Asia; northwards from the Mediterranean to the limits of tree-growth.
Stoat or Ermine (Mustela erminea, Linn.).
Though the gun and the snare of the gamekeeper and the poultry-farmer levy their toll upon the Stoat equally with the Polecat, and the keeper's gibbet always shows a goodly row of Stoats, the species manages to keep itself well represented, even in the strictly preserved woods of Southern England. There must, therefore, be some additional reason for the scarcity of the Polecat ([see p. 74]).
The Stoat is much smaller than the Polecat, its total length being only a little more than fourteen inches, of which about four and a half inches are the long-haired but not very bushy tail. In colour, too, it is very distinct, the upper parts being red-brown and the under surface white tinged with yellow. The tail takes the colour of the upper surface, except its tip which is invariably a tuft of long black hairs. In the Alpine districts of Scotland as in other northern countries, the fur in winter becomes pure white all over, with the exception of the tip of the tail which always remains black. This change takes place also in the North of England, but not so generally, and in the South it is only of rare occurrence, and often only partially, some parts remaining brown, as a ring around the eyes producing a spectacled appearance. The summer coloration is "protective" inasmuch that it harmonises generally with the colour of the ground littered with the remains of dead leaves, bark, etc.; but in a landscape under snow for months, as the Alpine districts are, the brown fur would render the animal so conspicuous that it would be heavily handicapped in the hunt for food; but the winter change to white fur enables the Stoat to steal upon its prey unseen from a short distance. The change is quite sudden, given the requisite fall in the temperature, the pigment being withdrawn. ([See Introductory chapter.])
Like the Polecat, the Stoat can secrete a most objectionable odour from its scent-glands, but in this case it is not nearly so insupportable. St. John says that if the Stoat is suddenly shot before he has had time to see his aggressor the dead body has not this offensive odour; the same result follows upon his sudden death in a spring trap, but if he is trapped alive or hunted before being shot the vile smell is imparted to the fur and is irremovable.
The Stoat hunts along hedgerows, rivers and brooks, in the latter places for fish, of which all the members of the Weasel tribe are exceedingly fond. An eel or other fish placed in a trap is a deadly bait for these animals. The Stoat also frequents sand dunes, where it lives sumptuously upon Rabbits. It is very destructive to game and poultry, which it will attack right in the open field, and if pursued by a dog, immediately takes shelter in a mole's or rat's run, where pursuit is impossible. It will destroy the Mole and take possession of its chamber, though it appears to be fonder of "field mice" (Voles) than of Moles. Although largely nocturnal in its habits, it is by no means exclusively so, and there is more chance of observing the Stoat hunting in broad daylight than in the case of any other of our native carnivora. Sometimes it hunts in small packs—family parties; and it is said that when through increase of its own numbers it has largely reduced the food supply of a district, it will migrate in large numbers, when their associated courage is so great that they will attack a man. A single female who has young will, indeed, exhibit the greatest courage and ferocity in their defence. The Stoat hunts by scent, and its movements consist largely of a succession of low bounds which give its progress a snake-like appearance—and like the other members of the family it makes sidelong leaps. Many years ago, whilst walking along a woodland road in Surrey, we paused to listen to cries of terror in the cover far ahead. A panic-stricken young Rabbit came into the open in our direction swiftly pursued by a Stoat which rapidly gained upon it. As it came near the Rabbit became aware of our presence and appeared deliberately to change its course, and fell on its side exhausted against our feet. The Stoat, by this time only a few yards away, stopped, and looked up at us with a snarling expression, but kept out of reach of our uplifted stick. Realising that the hunt had failed and the Rabbit had found a spoilsport protector, the Stoat then made off into the bracken; whilst the panting Rabbit allowed us to carry it on our arm for half a mile until it had recovered. Its natural fear of man was not nearly so great as its terror inspired by the bloodthirsty Stoat; and when at length it was set down in what was judged to be a safe place, it hopped off without any frightened haste.
It appears that the Hare under similar conditions does not exert itself greatly to escape from the Stoat, but becomes so terrorised as to be unable to adopt methods which so frequently outwit the Fox or the fleetness of trained hounds.
The nursery is made in a hole in the bank, the hollow of a decayed tree, or in the retreat of a female Mole who has been killed or evicted. Here about April or May the female Stoat gives birth to four or five young, which she will defend with great fierceness against all dangers.
The distribution of the Stoat extends eastward from Great Britain into Asia, and from the Alps and Pyrenees across Europe to its arctic shores.