The British flora is singularly poor in coniferous plants, the Scots Pine, the Yew, and the Juniper being our only native species, and even of these some authorities will have it that the Yew is not truly a Conifer at all; they place it in a separate order—Taxaceæ. For our present purpose, however, although the Yew does not produce cones, it will be convenient to keep it in its old position. The principal feature distinguishing all Conifers and their allies (Gymnosperms) from other flowering plants (Angiosperms) is briefly this: Angiosperms have their incipient seeds (ovules) enclosed in a carpel, through which a shoot from the pollen grain has to penetrate in order to reach and fertilize the ovule. In Gymnosperms the carpel takes the form of a leaf or bract, upon which the naked ovule lies open to actual
contact with the pollen grain. After fertilization the carpel enlarges to protect the seed, and becomes fleshy or woody, in the latter case a group of carpels forming the well-known cones of Pine or Fir.
In some of the groups (as the Yew, for example) the male or pollen-producing flowers are borne by a separate tree from that which bears the female or cone-producing flowers. In the Pines both sexes are found on the same tree; but throughout the order the pollen is carried by the wind. All the species are trees, or shrubs. They are among the most valuable of timber trees, and, in addition, yield a number of useful substances, such as pitch, tar, turpentine, etc. The leaves are always rigid, extremely narrow, and long in proportion: usually of the form that botanists term linear, with the two sides parallel. In the Yews these leaves spread out in two rows from opposite sides of the twigs; in the Pines they are in clusters of two, three, or five, seeming to be bound together at the base by a wisp of thin skin. The number of leaves in each bundle is often a help in distinguishing species.
The Yew (Taxus baccata) lacks the graceful proportions of most of our trees, but it has for compensation a most obvious air of strength and endurance. Who doubts, as he gazes at some sombre Yew in the old churchyard, the story of the local antiquarian, who tells him the tree has so stood for 2000 years. He may, perhaps, mildly suggest that neither the church nor the churchyard was in existence so far back, but even then the antiquarian will probably have the last word by suggesting that the grove of Yews of which this formed part was the church of the past. Thousands see in cathedral aisles the reproduction in stone of the pine-forest or the beech-wood. Standing before an ancient Yew they may see whence came the idea for those clustered columns. They actually exist in the bole of the Yew, which presents the appearance not of a single trunk, but of several trunks that have coalesced. This condition is due to the
Yew continually pushing out new shoots from the lower part of its bole, which take an upright direction, and coalesce with the old wood.
Although the Yew is a large tree, it is by no means a tall tree: the height of full-grown Yews in this country ranging between fifteen and fifty feet, though they are said to attain a greater length in India. The bole of the Yew is short but massive, covered with a thin red bark, that flakes off in patches much after the manner of Plane-bark. Large specimens have a girth of from twenty-five to fifty feet—or even more. Such a circumference represents the growth of many centuries, for the annual growth-rings are very thin. It is this very slow growth that produces the hard, compact, and elastic wood that was so highly esteemed in the days of the long-bow. Not only is the timber elastic and strong, but it is exceedingly durable, so that it is said, "A post of Yew will outlast a post of iron." Its branches start from the trunk at only a few feet from the ground, and, taking an almost horizontal direction, throw out a great number of leafy twigs, which provide a dense and extensive shade. These leaves are leathery in texture, curved somewhat after the manner of a reaping-hook, shiny and dark above, pale and unpolished below.
We have already mentioned that the Yew is a diœcious tree—that is, one whose male and female blossoms are borne on separate trees—but the statement requires qualification to this extent, that occasionally a tree will be found bearing a branch or branches whose flowers are of the sex opposite to those covering the greater part of the tree. The male catkin is almost round, a quarter of an inch across, and contains about half a dozen yellow anthers, the base surrounded by dry overlapping scales. They may be found during February and March, in profusion on the underside of the boughs. The female flower is much smaller, and consists of a fleshy disk with a few scales at its base, and on this stands a single seed-egg. After fertilization of the seed-egg