[fig 67]

[fig 68]


Fig. 65. Young Oceania just escaped from its reproductive calycle; magnified.

Fig. 66. The same as Fig. 65, from below, still more magnified; t long tentacles., t' rudimentary tentacle, e eye-speck on each side of base of tentacles.

Fig. 67. Young Oceania, older than Fig. 65; magnified.

Fig. 68. Adult Oceania; natural size.

Let us begin with its earlier condition. When it first escapes from the parent Hydroid stock, the Oceania is almost spherical in form. (See [Fig. 65.]) The disk is divided by four chymiferous tubes, running from the summit to the margin, where they meet the circular tube in which they all unite. At this time, it has but two well-developed tentacles, opposite each other on the margin of the disk, just at the base of two of the chymiferous tubes ([Fig. 66]), while two others are just discernible in a rudimentary state, forming slight projections at the base of the two other tubes. [Fig. 66] gives a view of the animal from below, at this stage of its growth, while [Fig. 65] shows it in profile. It will be seen by the latter how very spherical is the outline of the disk at this period, while the proboscis, in which are placed the mouth and digestive cavity, is quite long, and hangs down considerably below the lower surface of the disk. As the animal advances in age the disk loses its spherical outline, and becomes much flattened, as may be seen in [Fig. 67]. It may be well to introduce here some explanation of the law according to which the different sets of tentacles follow each other in successive cycles of growth, since it is a law of almost universal application in Jelly-fishes and Polyps; and, owing to the smaller number and simpler arrangement of the tentacles in Oceania, it may be more easily analyzed in them than in many others, where the number and complication of the different sets of tentacles make it very difficult to trace their relation to each other during their successive growth. We have seen that the Oceania begins life with only two tentacles. These form the first set, and are marked with the number 1 in the subjoined diagram, which gives the plan of all the different sets in their regular order. The second set, marked 2, consists also of two, which are developed at equal distances between the first two, i.e. at right angles with them. The third set, however, marked 3, consists of four, as do all the succeeding sets, and they are developed between the first and second. The fourth set comes in between the first and third; the fifth between the third and second; the sixth between the first and fourth; the seventh between the fifth and second; the eighth between the third and fourth; the ninth between the fifth and third. The ultimate number of tentacles in the Oceania is thirty-two, or sometimes thirty-six, and the cycles always in twos or multiples of two. But whatever be the number included in the successive sets of tentacles, and the unit for the first set ranges from two to forty-eight, the law in different kinds of Jelly-fishes is always the same, the youngest set always forming between the oldest preceding set. Thus the fourth set comes in between the first and third, and the fifth between the second and third, the intervals occupied now by the fourth set, being limited by the first set of tentacles on one side, and by the third set on the other side, while the intervals occupied by the fifth set are bounded by the second and third sets.

[fig 67b]

Diagram of succession of tentacles.

The little spheres represented between the tentacles on the margin of the disk, in Figs. [65-67], are eye-specks, and these continue to increase in number with age; in this the Oceania differs from the Eucope, in which it will be remembered there were but two eye-specks in each quarter-segment of the disk throughout life. [Fig. 68] represents the adult Oceania in full size, when it averages from an inch and a half to two inches in diameter. It is slow and languid in its movements, coming to the surface only in the hottest hours of the summer days; at such times it basks in the sun, turning lazily about, and dragging its tentacles after it with seeming effort. Sometimes it remains for hours suspended in the water, not moving even its tentacles, and offering a striking contrast to its former great activity when young, and to the lively little Eucope, which darts through the water at full speed, hardly stopping to rest for a moment. If the Oceania be disturbed it flattens its disk, and folds itself up somewhat in the shape of a bale (see [Fig. 69]), remaining perfectly still, with the tentacles stretching in every direction. When the cause of alarm is removed, it gently expands again, resuming its natural outline and indolent attitudes. The number of these animals is amazing. At certain seasons, when the weather is favorable, the surface of the sea may be covered with them, for several miles, so thickly that their disks touch each other. Thus they remain packed together in a dense mass, allowing themselves to be gently drifted along by the tide till the sun loses its intensity, when they retire to deeper waters. Some points, not yet observed, are still wanting to complete the history of this Jelly-fish. By comparing such facts, however, as are already collected respecting it, with our fuller knowledge of the same process of growth in the Eucope, Tima, and Melicertum, we may form a tolerably correct idea of its development. It is hatched from a Campanularia.

[fig 69]