Zoonomia; Or, the Laws of Organic Life, Vol. I

by Erasmus Darwin

In answer to this mode of reasoning it may be sufficient to observe, that the contractile fibres consist of inert matter, and when the sensorial power is withdrawn, as in death, they possess no power of motion at all, but remain in their last state, whether of contraction or relaxation, and must thence derive the whole of this property from the spirit of animation. At the same time it is not improbable, that the moving fibres of strong people may possess a capability of receiving or containing a greater quantity of the spirit of animation than those of weak people.

In every contraction of a fibre there is an expenditure of the sensorial power, or spirit of animation; and where the exertion of this sensorial power has been for some time increased, and the muscles or organs of sense have in consequence acted with greater energy, its propensity to activity is proportionally lessened; which is to be ascribed to the exhaustion or diminution of its quantity. On the contrary, where there has been less fibrous contraction than usual for a certain time, the sensorial power or spirit of animation becomes accumulated in the inactive part of the system. Hence vigour succeeds rest, and hence the propensity to action of all our organs of sense and muscles is in a state of perpetual fluctuation. The irritability for instance of the retina, that is, its quantity of sensorial power, varies every moment according to the brightness or obscurity of the object last beheld compared with the present one. The same occurs to our sense of heat, and to every part of our system, which is capable of being excited into action.

When this variation of the exertion of the sensorial power becomes much and permanently above or beneath the natural quantity, it becomes a disease. If the irritative motions be too great or too little, it shews that the stimulus of external things affect this sensorial power too violently or too inertly. If the sensitive motions be too great or too little, the cause arises from the deficient or exuberant quantity of sensation produced in consequence of the motions of the muscular fibres or organs of sense; if the voluntary actions are diseased the cause is to be looked for in the quantity of volition produced in consequence of the desire or aversion occasioned by the painful or pleasurable sensations above mentioned. And the diseases of associations probably depend on the greater or less quantity of the other three sensorial powers by which they were formed.

From whence it appears that the propensity to action, whether it be called irritability, sensibility, voluntarity, or associability, is only another mode of expression for the quantity of sensorial power residing in the organ to be excited. And that on the contrary the words inirritability and insensibility, together with inaptitude to voluntary and associate motions, are synonymous with deficiency of the quantity of sensorial power, or of the spirit of animation, residing in the organs to be excited.

[II]. Of sensorial Exertion.

[1]. There are three circumstances to be attended to in the production of animal motions, 1st. The stimulus. 2d. The sensorial power. 3d. The contractile fibre. 1st. A stimulus, external to the organ, originally induces into action the sensorial faculty termed irritation; this produces the contraction of the fibres, which, if it be perceived at all, introduces pleasure or pain; which in their active state are termed sensation; which is another sensorial faculty, and occasionally produces contraction of the fibres; this pleasure or pain is therefore to be considered as another stimulus, which may either act alone or in conjunction with the former faculty of the sensorium termed irritation.

This new stimulus of pleasure or pain either induces into action the sensorial faculty termed sensation, which then produces the contraction of the fibres; or it introduces desire or aversion, which excite into action another sensorial faculty, termed volition, and may therefore be considered as another stimulus, which either alone or in conjunction with one or both of the two former faculties of the sensorium produces the contraction of animal fibres. There is another sensorial power, that of association, which perpetually, in conjunction with one or more of the above, and frequently singly, produces the contraction of animal fibres, and which is itself excited into action by the previous motions of contracting fibres.

Now as the sensorial power, termed irritation, residing in any particular fibres, is excited into exertion by the stimulus of external bodies acting on those fibres; the sensorial power, termed sensation, residing in any particular fibres is excited into exertion by the stimulus of pleasure or pain acting on those fibres; the sensorial power, termed volition, residing in any particular fibres is excited into exertion by the stimulus of desire or aversion; and the sensorial power, termed association, residing in any particular fibres, is excited into action by the stimulus of other fibrous motions, which had frequently preceded them. The word stimulus may therefore be used without impropriety of language, for any of these four causes, which excite the four sensorial powers into exertion. For though the immediate cause of volition has generally been termed a motive; and that of irritation only has generally obtained the name of stimulus; yet as the immediate cause, which excites the sensorial powers of sensation, or of association into exertion, have obtained no general name, we shall use the word stimulus for them all.

Hence the quantity of motion produced in any particular part of the animal system will be as the quantity of stimulus and the quantity of sensorial power, or spirit of animation, residing in the contracting fibres. Where both these quantities are great, strength is produced, when that word is applied to the motions of animal bodies. Where either of them is deficient, weakness is produced, as applied to the motions of animal bodies.

Now as the sensorial power, or spirit of animation, is perpetually exhausted by the expenditure of it in fibrous contractions, and is perpetually renewed by the secretion or production of it in the brain and spinal marrow, the quantity of animal strength must be in a perpetual state of fluctuation on this account; and if to this be added the unceasing variation of all the four kinds of stimulus above described, which produce the exertions of the sensorial powers, the ceaseless vicissitude of animal strength becomes easily comprehended.