This dogmatic conception of the idea of species, and the arbitrary proceedings connected with it, necessarily led to the most perplexing contradictions, and to the most untenable suppositions. This is clearly demonstrable in the case of the celebrated Cuvier (born in 1769), who next to Linnæus has exercised the greatest influence on the study of zoology. In his conception and definition of the idea of species, he agreed on the whole with Linnæus, and shared also his belief in an independent creation of individual species. Cuvier considered their immutability of such importance that he was led to the foolish assertion—“The immutability of species is a necessary condition of the existence of scientific natural history.” As Linnæus’ definition of species did not satisfy him, he made an attempt to give a more exact and, for systematic practice, a more useful definition, in the following words: “All those individual animals and plants belong to one species which can be proved to be either descended from one another, or from common ancestors, or which are as similar to these as the latter are among themselves.”
In dealing with this matter, Cuvier reasoned in the following manner:—“In those organic individuals, of which we know that they are descended from one and the same common form of ancestors—in which, therefore, their common ancestry is empirically proved—there can be no doubt that they belong to one species, whether they differ much or little from one another, or whether they are almost alike or very unlike. Moreover, all those individuals also belong to this species which differ no more from the latter (those proved to be derived from a common stock) than these differ from one another.” In a closer examination of this definition of species given by Cuvier, it becomes at once evident that it is neither theoretically satisfactory nor practically applicable. Cuvier, with this definition, began to move in the same circle in which almost all subsequent definitions of species have moved, through the assumption of their immutability.
Considering the extraordinary authority which George Cuvier has gained in the science of organic nature, and in consequence of the almost unlimited supremacy which his views exercised in zoology, during the first half of our century, it seems appropriate here to examine his influence a little more closely. This is all the more necessary as we have to combat, in Cuvier, the most formidable opponent to the Theory of Descent and the monistic conception of nature.
One of the many and great merits of Cuvier is that he stands forth as the founder of Comparative Anatomy. While Linnæus established the distinction of species, genera, orders, and classes mostly upon external characters, and upon separate and easily discoverable signs in the number, size, place, and form of individual organic parts of the body, Cuvier penetrated much more deeply into the essence of organization. He demonstrated great and wide differences in the inner structure of animals, as the real foundation of a scientific knowledge and classification of them. He distinguished natural families in the classes of animals, and established his natural system of the animal kingdom on their comparative anatomy.
The progress from Linnæus’ artificial system to Cuvier’s natural system was exceedingly important. Linnæus had arranged all animals in a single series, which he divided into six classes, two classes of Invertebrate, and four classes of Vertebrate animals. He distinguished these artificially, according to the nature of their blood and heart. Cuvier, on the other hand, showed that in the animal kingdom there were four great natural divisions to be distinguished, which he termed Principal Forms, or General Plans, or Branches of the animal kingdom (Embranchments), namely—1. The Vertebrate animals (Vertebrata); 2. The Articulate animals (Articulata); 3. The Molluscous animals (Mollusca); and 4. The Radiate animals (Radiata). He further demonstrated that in each of these four branches a peculiar plan of structure or type was discernible, distinguishing each branch from the three others. In the Vertebrate animals it is distinctly expressed by the form of the skeleton, or bony framework, as also by the structure and position of the dorsal nerve-chord, apart from many other peculiarities. The Articulate animals are characterized by their ventral nerve-chord and their dorsal heart. In Molluscs the sack-shaped and non-articulate body is the distinguishing feature. The Radiate animals, finally, differ from the three other principal forms by their body being the combination of four or more main sections united in the form of radii (antimera).
The distinction of these four principal forms of animals, which has become extremely productive in the development of zoology, is commonly ascribed entirely to Cuvier. However, the same thought was expressed almost simultaneously, and independently of Cuvier, by Bär, one of the greatest naturalists, and still living, who did the most eminent service in the study of animal development. Bär showed that in the development of animals, also, four different main forms (or types) must be distinguished.[(20)] These correspond with the four plans of structure in animals, which Cuvier distinguished on the ground of comparative anatomy. Thus, for example, the individual development of all Vertebrate animals agrees, from the commencement, so much in its fundamental features that the germs or embryos of different Vertebrate animals (for example, of reptiles, birds, and mammals) in their earlier stages cannot be distinguished at all. It is only at a late stage of development that there gradually appear the more marked differences of form which separate those different classes and orders from one another. The plan of structure, which shows itself in the individual development of Articulate animals (insects, spiders, crabs), is from the beginning essentially the same in all Articulate animals, but different from that of all Vertebrate animals. The same holds good, with certain limitations, in Molluscous and Radiated animals.
Neither Bär, who arrived at the distinction of the four animal types or principal forms through the history of the individual development (Embryology), nor Cuvier, who arrived at the same conclusion by means of comparative anatomy, recognized the true cause of this difference. This is disclosed to us by the Theory of Descent. The wonderful and astonishing similarity in the inner organization and in the anatomical relations of structure, and the still more remarkable agreement in the embryonic development of all animals belonging to one and the same type (for example, to the branch of the Vertebrate animals), is explained in the simplest manner by the supposition of their common descent from a single primary original form. If this view is not accepted, then the complete agreement of the most different Vertebrate animals, in their inner structure and their manner of development, remains perfectly inexplicable. In fact it can only be explained by the law of inheritance.
Next to the comparative anatomy of animals and the systematic zoology founded anew by it, it was specially to the science of petrifactions, or Palæontology, that Cuvier rendered great service. We must draw special attention to this, because these very palæontological views, and the geological ideas connected with them, were held almost universally in the highest esteem during the first half of the present century, and caused the greatest hindrance to the working out of a truly natural history of creation.
Petrifactions, the scientific study of which Cuvier promoted at the beginning of our century in a most extensive manner, and established quite anew for the Vertebrate animals, play one of the most important parts in the “non-miraculous history of creation.” For these remains and impressions of extinct animals and plants, preserved to us in a petrified condition, are the true “monuments of the creation,” the infallible and indisputable records which fix the correct history of organisms upon an irrefragable foundation. All petrified or fossil remains and impressions tell us of the forms and structure of such animals and plants as are either the progenitors and ancestors of the present living organisms, or they are the representatives of extinct collateral lines, which, together with the present living organisms, branched off from a common stem.
These inestimable records of the history of creation throughout a long period played a subordinate part in science. Their true nature was indeed correctly understood, even more than five hundred years before Christ, by the great Greek philosopher, Xenophanes of Colophon, the same who founded the so-called Eleatic philosophy, and who was the first to demonstrate with convincing precision that all conceptions of personal gods result in more or less rude anthropomorphism.