When considering the struggle for life, we started from the fact that the number of germs which all animals and plants produce is infinitely greater than the number of individuals which actually come to life and remain alive for a longer or shorter time. Most organisms produce during life thousands or millions of germs, from each of which, under favourable circumstances, a new individual might arise. In most animals and plants these germs are eggs, that is cells, which for their development require sexual fructification. But among the Protista, the lowest organisms, which are neither animals nor plants, and which propagate themselves only in a non-sexual manner, the germ-cells, or spores, require no fructification. Now, in all cases the number of unsexual, as well as of sexual germs, is out of all proportion to the number of actually living individuals of every species.
Taken as a whole, the number of living animals and plants on our earth remains always about the same. The number of places in the economy of nature is limited, and in most parts of the earth’s surface these places are always approximately occupied. Certainly there occur everywhere and in every year fluctuations in the absolute and in the relative number of individuals of all species. However, taken as a whole, these fluctuations are of little importance, and it is broadly the fact that the total number of all individuals remains, on an average, almost constant. There is a constant fluctuation, which depends on the fact that in one year or another one or other series of animals and plants predominates, and that every year the struggle for life somewhat alters their relations.
Every single species of animals and plants would have densely peopled the whole earth’s surface in a short time, if it had not had to struggle against a number of enemies and hostile influences. Even Linnæus calculated that if an annual plant only produced two seeds (and there is not one which produces so few), it would have yielded in twenty years a million of individuals. Darwin has calculated of elephants, which of all animals seem the slowest to increase, that in seven hundred and fifty years the descendants of a single pair would amount to nineteen millions of individuals; this is supposing that every elephant, during its period of fertility (from the 30th to the 90th year), produced only three pairs of young ones, and survived itself to its hundredth year. In like manner the increase of the number of human beings—if calculated on the average proportion of births to population, and no hindrances to the natural increase stood in the way—would be such as to double the total in twenty-five years. In every century the total number of men would have increased sixteen-fold; whereas we know that the total number of human beings increases but slowly, and that the increase of population is very different in different countries. While European tribes spread over the whole globe, other tribes or species of men every year draw nearer to their complete extinction. This is the case especially with the redskins of America, and with the copper-coloured natives of Australia. Even if these races were to propagate more abundantly than the white Europeans, yet they would sooner or later succumb to the latter in the struggle for life. But of all human individuals, as of all other organisms, by far the majority perish at the earliest period of their lives. Of the immense quantity of germs which every species produce, only very few actually succeed in developing, and of these few it is again only a very small portion which attain to the age in which they can reproduce themselves (compare p. [161]).
From the disproportion between the immense excess of organic germs and the small number of chosen individuals which are actually able to continue in existence beside one another, there follows of necessity that universal struggle for life, that constant fight for existence, that perpetual competition for the necessaries of life, of which I gave a sketch in my seventh chapter. It is this struggle for life which brings natural selection into play, which in its turn is made use of by the interaction of the phenomena of Inheritance and Adaptation as a sifting agency, and which thus causes a continual change in all organic forms. In this struggle for acquiring the necessary conditions of existence, those individuals will always overpower their rivals who possess any individual privilege, any advantageous quality, of which their fellow competitors are destitute. It is true we are able only in the fewest cases (in those animals and plants best known to us) to form an approximate conception of the infinitely complicated interaction of the numerous circumstances, all of which here come into combination. Only think how infinitely varied and complicated are the relations of every single human being to the rest of mankind, and in general, to the whole of the surrounding outer world. But similar relations prevail also among all animals and plants which live together in one place. All influence one another actively or passively. Every animal and every plant struggles directly with a number of enemies, beasts of prey, parasitic animals, etc. Plants standing together struggle with one another for the space of ground requisite for their roots, for the necessary amount of light, air, moisture, etc. In like-manner, animals living together struggle with one another for their food, dwelling-place, etc. In this most active and complicated struggle, any personal superiority, however small, any individual advantage, may possibly decide the issue in favour of the one possessing it. This privileged individual remains the victor in the struggle, and propagates itself, while its fellow-competitors perish before they succeed in propagating themselves. The personal advantage which gave it the victory is transmitted by inheritance to its descendants, and by a further development may become so strongly marked as to cause us to consider the later generations as a new species.
The infinitely complicated correlations which exist between the organisms of every district, and which must be looked upon as the real conditions of the struggle for life, are mostly unknown to us, and are very difficult to discover. We have hitherto been able to trace them only to a certain point in individual cases, as in the example given by Darwin of the relations between cats and red clover in England. The red clover (Trifolium pratense), which in England is among the best fodder for cattle, requires the visit of humming-bees in order to attain the formation of seeds. These insects, while sucking the honey from the bottom of the flower, bring the pollen in contact with the stigma, and thus cause the fructification of the flower, which never takes place without it. Darwin has shown by experiments, that red clover which is not visited by humming-bees does not yield a single seed. The number of bees is determined by the number of their enemies, the most destructive of which are the field-mice. The more the field-mice predominate, the less the clover is fructified. The number of field-mice, again, is dependent upon the number of their enemies, principally cats. Hence in the neighbourhood of villages and towns, where many cats are kept, there are plenty of bees. A great number of cats, therefore, is evidently of great advantage for the fructification of clover. This example may be followed still further, as has been done by Carl Vogt, if we consider that cattle which feed on red clover are one of the most important foundations of the wealth of England. Englishmen preserve their bodily and mental powers chiefly by making excellent meat—roast beef and beefsteak—their principal food. The English owe the superiority of their brains and minds over those of other nations in a great measure to their excellent meat. But this is clearly indirectly dependent upon the cats, which pursue the mice. We may, with Huxley, even trace the chain of causes to those old maids who cherish and keep cats, and, consequently, are of the greatest importance to the fructification of the clover and to the prosperity of England. From this example we can see that the further it is traced the wider is the circle of action and of correlation. We can with certainty maintain that there exist a great number of such correlations in every plant and in every animal, only we are not always able to point out and survey their concatenation as in the last instance.
Another remarkable example of important correlations is the following, given by Darwin. In Paraguay, there are no wild oxen and horses, as in the neighbouring parts of South America, both north and south of Paraguay. This surprising circumstance is explained simply by the fact that in that country a kind of small fly is very frequent, and is in the habit of laying its eggs in the navel of newly-born calves and foals. The newly-born animals die in consequence of this attack, and the small deadly fly is therefore the cause of oxen and horses never becoming wild in that district. Supposing that this fly were destroyed by some insect-eating bird, then these large mammals would grow wild in Paraguay, as well as in the neighbouring parts of South America; and as they would eat a quantity of certain species of plants, the whole flora, and, consequently again, the whole fauna of the country would become changed. It is hardly necessary to state, that at the same time the whole economy, and consequently the character, of the human population would alter.
Thus the prosperity, nay, even the existence of whole populations can be indirectly determined by a single small animal or vegetable form in itself extremely insignificant. There are small coral islands whose human inhabitants live almost entirely upon the fruit of a species of palm. The fructification of this palm is principally effected by insects, which carry the pollen from the male to the female palm trees. The existence of these useful insects is endangered by insect-eating birds, which in their turn are pursued by birds of prey. The birds of prey, however, often succumb to the attack of a small parasitical mite, which develops itself in millions in their feathers. This small, dangerous parasite, again, may be killed by parasitical moulds. Moulds, birds of prey, and insects would in this case favour the prosperity of the palm, and consequently of man; birds, mites, and insect-eating birds would, on the other hand, endanger it.
Interesting examples in relation to the change of correlations in the struggle for life are furnished also by those isolated oceanic islands, uninhabited by man, on which at different times goats and pigs have been placed by navigators. These animals become wild, and having no enemies, they increase in number so excessively, that the rest of the animal and vegetable population suffer in consequence, and the island finally may become almost a waste, because there is insufficient food for the large mammals which increase too numerously. In some cases on an island thus overrun with goats and pigs, other navigators have let loose a couple of dogs, who enjoyed this superabundance of food, and they again increased so numerously, and made such havoc among the herds, that after several years the dogs themselves lacked food, and they also almost died out. The equilibrium of species continually changes in this manner in nature’s economy, accordingly as one or another species increases at the expense of the rest. In most cases the relations of different species of animals and plants to one another are much too complicated for us to be able to follow them, and I leave it to the reader to picture to himself what an infinitely complicated machinery is at work in every part of the world in consequence of this struggle. The impulses which started the struggle, and which altered and modified it in different places, are in the end seen to be the impulses of self-preservation—in fact, the instinct leading individuals to preserve themselves (the instinct of obtaining food), and the instinct leading them to preserve the species (instinct of propagation). It is these two fundamental instincts of organic self-preservation of which Schiller, the idealist (not Goethe, the realist!) says:
“Meanwhile, until philosophy Sustains the structure of the world, Her workings will be carried on By hunger and by love.”[4]
It is these two powerful fundamental instincts which, by their varying activity, produce such extraordinary differences in species through the struggle for life. They are the foundations of the phenomena of Inheritance and Adaptation. We have, in fact, traced all phenomena of Inheritance to propagation, all phenomena of Adaptation to nutrition, as the two wider classes of material phenomena to which they belong.