FIG. 4.—The cartilages of the larynx or voice-box. A large portion of the shield cartilage on the right side has been cut away, in order to show the two pyramid cartilages; these are seen jointed by their bases with the ring cartilage; anteriorly are seen the two vocal processes which give attachment to the two vocal cords (white ligaments), which extend across the voice-box to be inserted in front in the angle of the shield cartilage. Groups of muscles pull upon these cartilages in such a manner as to increase, or diminish, the chink between the vocal cord in ordinary inspiration and expiration; in phonation a group of muscles approximate the cords, while another muscle makes them tense.
The Larynx.—The larynx is situated at the top of the sound-pipe (trachea or windpipe), and consists of a framework of cartilages articulated or jointed with one another so as to permit of movement (vide [fig. 4]). The cartilages are called by names which indicate their form and shape: (1) shield or thyroid, (2) the ring or cricoid, and (3) a pair of pyramidal or arytenoid cartilages. Besides these there is the epiglottis, which from its situation above the glottis acts more or less as a lid. The shield cartilage is attached by ligaments and muscles to the bone (hyoid) in the root of the tongue, a pair of muscles also connect this cartilage with the sternum or breastbone. The ring cartilage is attached to the windpipe by its lower border; by its upper border in front it is connected with the inner surface of the shield cartilage by a ligament; it is also jointed on either side with the shield cartilage. The posterior part of the ring cartilage is much wider than the anterior portion, and seated upon its upper and posterior rim and articulated with [!-- pagenumber --]it by separate joints are the two pyramidal cartilages (vide [fig. 4]). The two vocal cords as shown in the diagram are attached to the shield cartilage in front, their attachments being close together; posteriorly they are attached to the pyramidal cartilages. It is necessary, however, to describe a little more fully these attachments. Extending forwards from the base of the pyramids are processes termed the "vocal processes," and these processes give attachment to the elastic fibres of which the vocal cords mainly consist. There are certain groups of muscles which by their attachment to the cartilages of the larynx and their action on the joints are able to separate the vocal cords or approximate them; these are termed respectively abductor and adductor [!-- pagenumber --]muscles (figs. [5] and [6]). In normal respiration the posterior ring-pyramidal muscles contract synergically with the muscles of inspiration and by separating the vocal cords open wide the glottis, whereby there is a free entrance of air to the windpipe; during expiration this muscle ceases to contract and the aperture of the glottis becomes narrower (vide [fig. 10]). But when the pressure is required to be raised in the air passages, as in the simple reflex act of coughing or in vocalisation, the glottis must be closed by approximation of the vocal cords, and this is effected by a group of muscles termed the adductors, which pull on the pyramid cartilages in such a way that the vocal processes are drawn towards one another in the manner shown in [fig. 7]. Besides the abductor and adductor groups of muscles, there is a muscle which acts in conjunction with the adductor group, and by its attachments to the shield cartilage above and the ring cartilage below makes tense the vocal cords (vide [fig. 5]); it is of interest to note that this muscle has a separate nerve supply to that of the abductor and adductor muscles.
Fig. 5
Diagram after Testut (modified), showing the larynx from the front.
Fig. 6
Diagram after Testut (modified), showing the posterior view of the larynx with the muscles.
On the top of the pyramid cartilages, in the folds of mucous membrane which cover the whole inside of the larynx are [!-- pagenumber --]two little pieces of yellow elastic cartilage; and in the folds of mucous membrane uniting these cartilages with the leaf-like lid cartilage (epiglottis) is a thin sheet of muscle fibres which acts in conjunction with the fibres between the two pyramid cartilages (vide [fig. 8]). I must also direct especial attention to a muscle belonging to the adductor group, which has another important function especially related to vocalisation: it is sometimes called the vocal muscle; it runs from the pyramid cartilage to the shield cartilage; it apparently consists of two portions, an external, which acts with the lateral ring-shield muscle and helps to approximate the vocal cords; and another portion situated within the vocal cord itself, which by contracting shortens the vocal cord and probably allows only the free edge to vibrate; moreover, when not contracting, by virtue of the perfect elasticity of muscle the whole thickness of the cord, including this vocal muscle, can be stretched and thrown into vibration (vide [fig. 8]). In the production of chest notes the whole vocal cord is vibrating, the difference in the pitch depending upon the tension produced by the contraction of the tensor (ring-shield) muscle. When, however, the [!-- pagenumber --]change from the lower to the upper register occurs, as the photographs taken by Dr. French and reproduced in a lecture at the Royal Institution by Sir Felix Semon show, the vocal cords become shorter, thicker, and rounder; and this can be explained by supposing that the inner portion of the vocal muscle contracts at the break from the lower to the upper register (vide [fig. 11]); and that as a result only the free edges of the cords vibrate, causing a change in the quality of the tone. As the scale is ascended the photographs show that the cords become longer and tenser, which we may presume is due to the continued action of the tensor muscle. Another explanation is possible, viz. that in the lower register the two edges of the vocal cords are comparatively thick strings. When the break occurs, owing to the contraction of the inner portion of the vocal muscle, we have a transformation into thin strings, at first short, but as the pitch of the note rises, the thin string formed by the edge of the vocal cord is stretched and made longer by the tensor. It should [!-- pagenumber --]be mentioned that Aikin and many other good authorities do not hold this view.