form; there appears to be but a single species, though more than one name has been given to supposed distinct species. As already mentioned, it differs from the Manatee in the possession of a Whale-like tail; the nostrils, too, are more upon the upper surface of the head, and there are no nails upon the flipper. The peculiar cleft lip of the Manatee is not so well developed in the Dugong, but there are traces of it; and in the foetus the likeness to the Manatee in this respect is very striking. It would thus appear that Halicore is a stage in advance upon Manatus; that the remarkable mechanism of the lip of the latter has been possessed, but has been lost, by the Dugong. The skull of the Dugong is distinguished by the stout premaxillary bones, which bear a tusk in the male. In the female the tooth is there, but is lodged within the bone. This incisor has a milk forerunner. The back teeth of the Dugong (there are no canines) are few in number (four or five, even six), thus showing a gradual reduction when compared with Manatus; and this culminates in the toothless Rhytina. It is also interesting to notice that in the massive lower jaw there are traces of an incisor. Were this to be developed into a tusk, the jaw would present a curious resemblance to that of Dinotherium.
The Dugong, H. dugong, has the reputation of being the original of the mermaid legends, since the young is held to the pectorally-situated breast with one flipper. "But it should be remembered," justly observes Dr. Blanford, "that stories of beings half man or woman, half fish, are as common in temperate as in tropical seas, and that some of them are more ancient than any European knowledge of the Dugong."
Extinct Sirenians.—The earliest genus that can be with certainty referred to this order is the Oligocene Prorastoma. This genus, though offering no particular skull-characters that assist in the determination of the much-debated affinities of the Sirenia, shows a remarkable condition of the teeth that may afford a clue. The species P. veronense, recently described by Mr. Lydekker,[[220]] is founded upon a fragment of the skull which contains two teeth apparently representing the third and fourth upper milk molars. The interest attaching to these teeth lies in the fact that they clearly exhibit the buno-selenodont condition characteristic of certain early Artiodactyles, e.g. Merycopotamus.
Halitherium is a later genus, which is known by the nearly
complete skeleton. The skull is like that of other Sirenia, with the down-turned premaxillary region. But the nasal bones, lost, or at least rudimentary, in recent forms, are well developed; the likeness of ancient to living forms in this respect being exactly paralleled by the Zeuglodonts, when compared with recent Whales. The vertebral centra exhibit distinct epiphyses, which have disappeared in living Sirenians. The cervical vertebrae are seven, of which the second and third are occasionally fused. There are nineteen pairs of ribs, and there are three lumbar vertebrae. The sternum consists of three separate pieces. There is a rudimentary femur.
The recently-extinct Steller's Sea-cow, belonging to the genus Rhytina, was a huge beast, seen in the flesh up to nearly the end of the last century. It frequented the shores of Bering's Straits. Its remains occur in the peat on the shores of those seas. It reaches a length of some 20 to 30 feet. The external characters were much like those of other recent Sirenians. The nostrils were above the fore part of the snout, the latter being truncated and obtuse. The tail was of the Cetacean pattern, and thus like that of Halicore. The head of this Sirenian was small, and the teeth had entirely vanished save for the apparent existence as transitory structures of two small incisors in the upper jaw. The absence of teeth was compensated by the presence of a horny palate for the trituration of the sea-weeds which constituted the food of Steller's Sea-cow. The fore-limbs seem to have possessed no nails, but were covered at the extremity with short, bristly hairs, no doubt serving the purpose of keeping the animal moored in safety to the slippery beds of Fucus upon which it browsed.
There are nineteen pairs of ribs. The vertebrae of the cervical region are the customary seven, and the centra are thin and plate-like as in the Cetacea, the animal being thus short-necked like those marine creatures.