Fig. 198.—Ocelot. Felis pardalis. × 1⁄10.
The Puma, F. concolor, the American Lion as it is called in the north, is a rather smaller animal than the last, and of a uniform tawny colour, tending to white on the abdomen and to a dark stripe along the back. The young, as already mentioned, are very distinctly spotted. Like the Tiger, the Puma can endure extremes of heat and cold; it is equally at home in the snow of North America and among the tropical forests and swamps of the south. It is a ferocious creature so far as concerns Deer, Lamas, Raccoons, even Skunks and Rheas, but, according to Mr. W. H. Hudson, will not attack man, and will even defend him against the Jaguar.[[264]] In captivity the Puma will purr like a Cat.
The Eyra, F. eyra, is another self-coloured American cat, which has a curious likeness to the totally distinct Cryptoprocta of Madagascar.
The Wild Cat of Europe, F. catus, is found over the greater part of Europe, and also in Northern Asia. It was undoubtedly common at one time in this country, though it appears never to have extended its range into Ireland. But the real Wild Cat is now rare in this island, and is confined to certain districts in
Scotland. Plenty of alleged wild Cats have been seen and even shot; but these are too frequently merely feral Cats, i.e. domestic tabbies which have reverted to a hunting life. The real Wild Cat differs from the domestic races by the proportionately longer body and limbs, the shorter and thicker tail; the pads of the toes are not quite black. The period of the gestation of the Wild Cat, according to Mr. Cocks, is a week or so longer than that of any domestic Cat.
The Domestic Cat is in fact regarded as the descendant of the Eastern F. caffra, or (perhaps and) the closely-allied F. maniculata. It is highly probable, however, that after introduction into this country as a domestic animal it has interbred with the Wild Cat. Many allied species of Cats will interbreed, even two so far apart as the Lion and the Tiger. There are interesting archaeological and linguistic reasons for regarding the Domestic Cat as an importation. The legend of Dick Whittington's Cat points to it being a rare and valuable animal, which a tamed F. catus would not at that time have been. There was an enactment in Wales of a penalty against him who should kill the king's Cat, again suggestive of its rarity and consequent value. The very name "Puss" is a hint of a foreign origin. Some would derive it from Perse, and upon this is based the notion that the Cat is from Persia. But it seems that Puss is the same as Pasht and Bubastis, showing so far an Egyptian origin for the animal. The ancestral Cats mentioned above are natives of Egypt.[[265]]
The genus Cynaelurus, which includes but a single species, C. jubatus, the Cheetah or Hunting Leopard, is separated from Felis by a number of characters. In the first place the claws are non-retractile, or at least less retractile than those of the true Cats. It is, moreover, longer legged. The molar is more in a line with the other teeth of the jaw, and the upper carnassial tooth has no inner tubercle. Messrs. Windle and Parsons have lately pointed out many Dog-like features in the muscles. This animal is about as large as a Leopard, but has plain black spots. As its vernacular name implies it is used for sport, and is quite easily tameable. It will purr like the Puma. The Cheetah occurs in India, Persia, Turkestan, and also in Africa; the latter
form is sometimes, though quite unnecessarily, separated as C. lanigera. The genus occurs fossil in the Siwalik deposits of India, the species being known as C. brachygnatha.
Fam. 2. Machaerodontidae.—This is a family of totally-extinct Cats which range from the Eocene down to the Pleistocene. Their general structure is like that of the Felidae; but they differ in a number of skeletal features. Thus there is an alisphenoid canal, and, as in Bears, there is a postglenoid foramen. There is also a distinct carotid foramen, which does not occur in the true Cats. The teeth are often distinguished by the huge size of the superior canines, which are "weapons for penetrating wounds, without rival among carnivorous animals." These must have been displayed at the sides of the chin when the mouth was closed, and it has even been suggested that the animal possessing these exaggerated canines could hardly have properly closed its mouth. The lower canines were often on the contrary much reduced, and in fact incisor-like. In tracing the series of these Cats we find a gradual reduction of the teeth from a more nearly complete number down to the specialised dentition of the existing Cats. The genus Proaelurus, Miocene in range, had four premolars in each jaw, and two molars in the lower and one in the upper. This is the greatest number of teeth found in any member of the group.
The resemblance of this genus to Cryptoprocta has been insisted upon. Archaelurus has suffered a reduction, since one premolar in the lower jaw has disappeared, its formula being thus I 3/3 C 1/1 Pm 4/3 M 1/2. The next stage is shown by Dinictis with three premolars in both jaws. There are a good many species of this genus which are all American and Miocene. This genus has five toes upon the hind-feet, and was probably plantigrade. It had retractile claws.