A variable number of the anterior cheek teeth may be little more than simple conical teeth; but the rest of the set are commonly more complicated. No definite laws can be laid down as to the complication of the posterior as compared with the anterior set. Broadly speaking, it is purely herbivorous creatures in which the least difference can be detected at the two extremities, and which are at the same time the most elaborately decorated with tubercles and ridges. The converse is true that in purely carnivorous animals, including insect- and fish-eating forms, there is the greatest difference between the anterior set of grinding teeth and those which follow. In these two respects such animals as a Lemur and a Rhinoceros occupy the extremes. Furthermore, it may be said that omnivorous creatures lie, as their diet would suggest, in an intermediate position. Generally speaking, when there is a marked difference between the first premolar and molars at the end of the series, there is a gradual approximation in structure of a progressive kind. The tubercles become more numerous in successive teeth; but the corollary which is apparently deducible from this, i.e. that the last molar is the most elaborate of the series, is by no means always true. The last cheek tooth indeed is often degenerate. On the other hand, it is very markedly the largest of the series in such diverse types as the Elephant, the hog Phacochoerus, and the Rodent Hydrochoerus. It is a rule that the cheek teeth of the upper jaw are more complicated than the corresponding teeth of the lower jaw.
The structure of the cheek teeth is very diverse among the Mammalia. Broadly, two types are to be recognised. There are
teeth in which the grinding surface is raised into a series of two, to many, tubercles sharper or blunter as the case may be;—sharper and fewer at the same time in carnivorous and especially in insectivorous types, more abundant in omnivorous animals. To this form of tooth the term "bunodont" is applied. There is no doubt that this is the earliest type of tooth; but whether the fewer or the more cusped condition is the primitive one is a question that is reserved for consideration at the end of the present chapter. The other type of grinding tooth is known as "lophodont." This is exemplified by such types as the Perissodactyla and Ungulates generally, and by the Rodents. The tooth is traversed by ridges which have generally a transverse direction to the long axis of the jaw in which the tooth lies. The ridges may be regarded as having been developed between tubercles which they connect and whose distinctness as tubercles is thereby destroyed. Lophodont teeth are only found in vegetable-feeding animals.
Fig. 36.—Molar teeth of Aceratherium platycephalum. × ½. m.1-m.3., Molars; mh, metaloph; p.1-p.4, premolars; ph, protoloph; ps.f, parastyle fossa; te, tetartocone. (After Osborn.)
The special characteristics of the teeth of various groups of animals will be considered further under the accounts of the several orders of recent and fossil Mammalia.
Fig. 37.—Two stages in the development of the teeth of a Mammal (diagrammatic sections). alv, Bone of alveolus; dent, dentine; dent.s, dental sac; en, enamel; en.m, enamel membrane; en.m2, enamel membrane of permanent tooth; en.plp, enamel pulp; gr, dental groove; lam, dental lamina; lam′, part of dental lamina which grows downwards below the tooth germ; n, neck connecting germs of milk and permanent tooth; pap, dental papilla; pap2, dental papilla of permanent tooth. (After O. Hertwig.)
A very general feature of the teeth of the Mammalia is what is usually termed the diphyodont dentition. In the majority of cases there are two sets of teeth developed, of which the first lasts for a comparatively short time, and is termed on account of its usual time of appearance the "milk dentition"; this is replaced later by the permanent dentition. In lower vertebrates the teeth are replaced as worn away. There is not, however, so great an antithesis in this matter between the Mammalia
and other vertebrates as was at one time assumed. But in order to explain this very important part of the subject it will be necessary to give some account of the development of the teeth. The type selected is the Hedgehog, which has been recently and carefully described by Dr. Leche of Stockholm, which type has furthermore the advantage of being a "central" type of mammal. The first step in the formation of the teeth is a continuous invagination of the epithelium covering the jaw to form a deepish wall of tissue running in the thickness of the jaw; this is perfectly continuous from end to end of the lower jaw. From this "common enamel germ" (Schmelzleiste of the Germans[[24]]) "special enamel germs" (Schmelzorgane, enamel organs) are developed here and there as thickenings in the form of buds