The Ring-tailed Phalangers, Pseudochirus, are more widely distributed than the last two genera; they range from Tasmania in the south to New Guinea in the north. They are not, however, ring-tailed, though the tip of the tail is generally white. As in the last genera, which have prehensile tails, the end of this appendage is naked. The mammae are four. The tooth formula is I 3/2 C 1/0 Pm 3/3 M 4/4. There are some ten species of the genus.
The Striped Phalanger, Dactylopsila trivirgata, is an animal about a foot long, whose identity can be ascertained by its striped, black and white skin. It is an arboreal creature that lives apparently both on leaves and grubs like so many arboreal creatures of quite different groups—Squirrels, for instance, and New-World Monkeys. The tooth formula is I 3/3 C 1/6 Pm 3/2 M 4/4.
Gymnobelideus leadbeateri is a small creature with a body 6 inches in length. It is restricted to the colony of Victoria. The general look is that of Petaurus; the ears are naked.
Dromicia is a genus of Phalangers which although devoid of a parachute, such as is possessed by certain genera that will be considered immediately, is able to leap with great agility from branch to branch. The ears are large and thin and almost naked; the tooth formula is I 3/2 C 1/0 Pm 3/3 M 4/4. They are minute creatures, the longest measuring, with the tail, but 10 inches. Dormouse-Phalanger is a name sometimes given to them. There are four species, ranging from Tasmania to New Guinea. The name Dormouse as applied to the genus seems to be owing to the way in which they hold a nut in the paws when feeding. D. nana is 4 inches long, with a tail of nearly the same length. It is thick at the base.
Distaechurus is the last genus of non-flying Phalangers. Its name refers to the arrangement of the hairs on the tail, which are disposed on either side in a row like the vane of a feather. The tooth formula is I 3/2 C 1/0 Pm 3/2 M 3/3, very nearly as in Acrobates. The ears are as in that genus.
Petaurus is the first genus of the Flying Phalangers, all of which are provided with a parachute-like expansion of the skin between the fore- and hind-limbs; the ears are large and naked; and the tooth formula is I 3/2 C 1/0 Pm 3/3 M 4/4. There are three
species of the genus, which extend through pretty well the entire Australian region. The term "flying" as applied to these and the other "flying" genera is of course an exaggeration. The animals cannot fly upwards; they can only descend in a skimming fashion, the folds of skin breaking their fall. P. breviceps is perhaps the best-known species. The body is 8, the tail 9 inches long.
Petauroides seems to be chiefly distinguished from Petaurus by the fact that, as in its ally Dactylopsila, the tail is partly naked terminally. In Petaurus and Gymnobelideus the tail is bushy to the very end, including its extreme tip below.
A third genus of Flying Phalangers is the minute Acrobates, which has a distichous tail like that of Distaechurus. It is not more than 6 inches in length including the tail. As to these Flying Phalangers it is exceedingly instructive to observe that the same method of "flight" has been apparently evolved three times; for the three genera are each of them specially related to a separate type of non-flying Phalanger. The same observation can be made about the Flying Squirrels, Anomalurus and Sciuropterus. The dental formula is I 3/2 C 1/0 Pm 3/3 M 3/3. The ears are thinly clad with hair. There are four teats.
Sub-Fam. 2. Phascolarctinae.—The Koala, or Native Bear, Phascolarctos cinereus, is the only representative of its sub-family. It is, like the Wombat, aberrant in the lack of an obvious tail. The absence of this appendage is curious in an arboreal creature whose near allies have a long and prehensile one. The structure of the Koala was investigated by the late Mr. W. A. Forbes.[[83]] There are some unexpected points of likeness to the Wombat: thus they agree in the absence of the tail, in the structure of the stomach, and in the great subdivision of the lobes of the liver. The brain, however, is smooth, and the caecum is exceedingly large and complicated in structure, that of the Wombat being short. That both animals have cheek-pouches is perhaps due to similar habits of temporarily storing masses of food. This animal has only eleven pairs of ribs. The tail has only seven or eight vertebra, and these have no chevron-bones.