Fig. 71.—Wombat. Phascolomys wombat. × 1⁄12.
Sub-Fam. 3. Phascolomyinae.—Phascolomys, the Wombat, is the only genus of this sub-family. This animal has the appearance of a heavily-built Marmot, like which it has a mere stump for a tail, and a pair of strong chisel-shaped and Rodent-like incisors, which, however, differ from those of Rodents in having a complete coating of cement. All the teeth of the animal are rootless, and there are no canines. The incisors have enamel on the front and lateral faces only. The dental formula is I 1/1 C 0/0 Pm 1/1 M 4/4. The affinities with other Diprotodont Marsupials are shown by the commencing syndactyly of the second and third toes. The rhinarium is naked or hairy. There is a rudimentary cheek-pouch, as in Phascolarctos. The Wombat has, like the Koala, and also the Beaver—which does away with some of the value of the comparison—a peculiar gland-patch in the stomach, a raised area of collected glands. In no other Marsupial is such a structure found, "whilst in the two forms under consideration its identity is almost precise. That such a unique structure should have been independently developed in two forms unrelated to each other, appears to me to be in the highest degree improbable." This is Mr. Forbes' opinion. It might be strengthened by adding the observation that, as there are other points of likeness between the Wombat and the Koala, it seems more unlikely that a structure so nearly identical should have been twice
developed in two not very distant forms. As in the Kangaroos, the atlas is open below. Ph. ursinus has 15 ribs; the other species the normal (for Marsupials) 13. Other points of likeness will be mentioned under the description of the Koala. These animals mainly feed upon roots; they live in companies in burrows. There are three species—Ph. ursinus, Ph. latifrons, and Ph. mitchelli. Ph. ursinus is Tasmanian in range, the other two species South Australian.
Fig. 72.—Skull of Wombat. Phascolomys wombat. (Lateral view.) ang, Angular process; cond, condyle of mandible; ex.oc, exoccipital; ext.aud, opening of bony auditory meatus; ju, jugal; lcr, lachrymal; max, maxilla; nas, nasal; p.max, premaxilla; sq, squamosal; ty, tympanic. (From Parker and Haswell's Zoology.)
Sub-Fam. 4. Tarsipedinae.—The genus Tarsipes ought perhaps to be removed from the present family. There is but a single species, which is a small creature of 7 inches in total length, of which the tail measures 4 inches. The teeth are much dwindled, the formula being I 2/1 C 1/0 Pm 1/0 M 3/3 = 22. The lower incisors are procumbent. The lower jaw, moreover, has not the characteristic Marsupial inflection. The intestinal canal is without the caecum present in the remaining Phalangeridae. It is a curious fact that this aberrant little Phalanger should come from Western Australia, like the even more aberrant Myrmecobius. Like the latter also, Tarsipes has a long exsertile tongue, with which, however, it extracts honey from flowers. Probably it also catches minute insects in the corollas of the flowers. It has been proved, in fact, that in captivity at any rate the animal is insectivorous; for it has been known to eat moths.
Fam. 3. Epanorthidae.—The extinct Epanorthidae of
Patagonia are represented to-day by a small Marsupial which has been rediscovered within the last two or three years. This little animal, formerly called Hyracodon (a pre-occupied name), is now termed Caenolestes, and is a native of Colombia and Ecuador. There are two species, and of these C. obscurus is called by the inhabitants "Raton runcho," which means opossum-rat. It lives apparently upon bird's eggs and small birds, though it belongs to the Diprotodont division of the Marsupials. Caenolestes, however, although diprotodont, has not the syndactylous character of the digits of the feet already referred to in the Kangaroos and their allies. The pouch is small and rudimentary. The dentition is I 4/3 C 1/1 Pm 3/3 M 4/4 = 46, and the teeth are said by Mr. Thomas to be much like those of the Australian Dromicia.[[84]]
In the skull a peculiarity which does not bear upon its affinities to other Marsupials, but is still interesting, is mentioned by Mr. Thomas. The nasals are not sufficiently prolonged to meet the upper edge of the maxillae, and so a vacuity is left, as in the skulls of many Ruminants (e.g. the Sable Antelope). The palate is very imperfect; the foramina, which render it so, reach as far forward as the last premolar. The lower jaw has quite the appearance of that of a Macropus or Phalanger, with long and forwardly projecting incisors.
Extinct Diprotodonts.—The great Diprotodon is a creature with a skull a yard long, which must have been of the size of a large Rhinoceros. Though closely allied to Macropus, it seems that this great beast did not hop after the fashion of a Kangaroo, its limbs being of a more equal size than in the Kangaroo. Recently some further remains of Diprotodon have been discovered in a lake known as Lake Mulligan, where they had apparently been bogged. Professor Stirling has contributed an account of these remains, which fills up a considerable gap in our knowledge. He has been able to state the structure of the fore- and hind-limbs. Both limbs are pentadactyle, the fingers of the fore-limb being approximately equal in length and general development. In the hind-limb the hallux is small, and consists of the metatarsal only. This bone is fixed in the position of "extreme abduction," and is suggestive of an arboreal limb. Digits two and three may have