Fig. 81.—Dasyure. Dasyurus viverrinus. × 1⁄5. (After Vogt and Specht.)
The next genus of this family, Dasyurus, comprises five species, which range over the whole of the Papuan and Australian sub-regions. The general form is Viverrine, and the hallux is sometimes present though small. The dental formula is as in the last genus, but the teeth "are more insectivorous in their character." There are six or eight mammae. The members of this genus are grey or brown, and spotted with white; they are all arboreal, and feed largely upon birds and their eggs. Mr. Thomas has pointed out that in two species, D. viverrinus and D. geoffroyi, the striae upon the foot-pads are absent, and that therefore these at least are probably not so purely arboreal as the rest. The animals are not diurnal, and during the day hide themselves in the hollow trunks of trees. They are spoken of as "Native Cats," but have the general habits of Martens. D. maculatus is common in Tasmania, but is rare in Australia, thus "approaching the condition now exhibited by the Thylacine and
Tasmanian Devil, namely, complete extermination in Australia, where both once lived." D. hallucatus shows an approach to Phascologale in its five-toed hind-feet and slender build.
Phascologale is a genus which, like the last, is usually arboreal (although not P. virginiae of North Queensland), but is of much smaller size, the species not exceeding the dimensions of a rat. They have no spots, but there is sometimes a stripe down the back. There are thirteen species, which have the same range as the last genus. The hallux is present though small, but the pouch is "practically obsolete," though there is a small fold of skin behind the teats. The rhinarium is naked; the tail is long, "bushy, crested, or nearly naked." The mammae are four to ten in number. The dental formula is as in Dasyurus, and the teeth are not very different in form; sometimes the last premolar is wanting. "The members of this genus," remarks Mr. Thomas, "evidently take the place in the Australian region filled in the Oriental by the Tupaiae, and in the Neotropical by the smaller Opossums."
The genus Sminthopsis comprises not more than four species, even smaller than the last. The largest species, S. virginiae, is only 125 mm. in length. The hallux is present, and there is a well-developed pouch. There are forty-six teeth, as in the Dasyures. The feet are narrow with granulated or hairy soles, whereas in Phascologale they are broad with smooth soles. The mammae are eight or ten. The genus ranges through Australia and Tasmania.
The genus Antechinomys has but a single species, which is a native of Queensland and New South Wales. The build is Jerboa-like, and the animal is, as might be inferred, terrestrial. The ears are very long, and the limbs elongated; the hallux is absent; the teeth are exactly as in Sminthopsis.
Antechinomys has thirteen dorsal and seven lumbar vertebrae; three sacrals and twenty-five caudals, the latter number being in excess of that of its allies. The stomach is nearly globular, with approximated orifices; the intestine measured 6.8 inches, a little more than twice the length of the animal itself. A. lanigera is a native of East Central Australia, and appears to be entirely terrestrial in habit, and to progress by a series of leaps—at any rate when going at full speed.
Professor Spencer, who found examples of this rare species, gives
an interesting description of its habits. Antechinomys has much the look of the Australian Rat, Hapalotis mitchelli; and as the two animals lead a similar kind of life, the resemblance is not unexpected. Professor Spencer wonders why these creatures are saltatory in habit. The country which they inhabit is arid, but with patches of grass and shrubs. For a big kangaroo the advantage of the power of leaping over such obstacles may be obvious, but not for the small and slender Antechinomys. The chief foes of this rare Marsupial appear to be predatory birds; and Professor Spencer thinks that the saltatory mode of progression may be more baffling to such pursuers than even a rapid run.
The genus Dasyuroides has been lately instituted by Professor Spencer for a Marsupial from Central Australia somewhat intermediate between Sminthopsis and Phascologale. As there is but one species, the generic will be considered with the specific characters. D. byrnei is an animal of about the size of the Common Rat. The hallux is absent. The tail is fairly thick, but not "incrassated." There are six mammae, and the pouch is but slightly developed, with two low lateral folds. The dentition is I 4/3 C 1/1 Pm 3/2 M 4/4. This Marsupial is nocturnal, and burrowing in habit. Its food consists of insects.[[87]]