The significance of these phenomena lies in the fact that in the evolutionary series the abnormal occurrences, which are found in different races in varying frequency, appear as new developments, which, if they should become normal, would increase the differentiation between man and the lower forms. The actual proof of increasing frequency of these features, and of their becoming permanent characteristics, has not been given, but seems more than plausible.
This inference is strongly supported by the occurrence of rudimentary, functionless organs, and by the temporary appearance of lower features during ontogenetic development.
It has been shown that some of these reductions—like the retention of outer incisors—are hereditary, and thus liable to perpetuate themselves. To a certain extent this explains the observation that certain variations occur with greater frequency among primitive tribes than in civilized man. Most primitive tribes are very small in numbers, or for long periods, during which they increased in numbers, have had little intercourse with foreign people. If in such a group any of the original families showed a certain peculiarity, it must now be found more frequently than in any other tribes. A case of this kind is the frequency of supernumerary vertebræ among the Indians of Vancouver Island, and probably also the frequency of the torus palatinus among the Lapps. It may be left an open question, whether the frequent occurrence of the os Incæ among the Pueblo Indians and Peruvians may be explained by the same consideration. Therefore it may be that the greater variability of certain races, in regard to these phenomena, is not an expression of a lower or higher degree of development of the whole group, as the case may be, but of the presence of a great number of members of a family which possessed the peculiar character. We do not deal in these cases with spontaneous variations, but with their hereditary re-appearance. In other words, if we are to admit the conclusion that greater variability means a lower or higher stage of development, it will be necessary first to prove that these variations appear spontaneously in any member of the group, and do not belong to certain families in which the feature is hereditary. Otherwise it will be necessary to prove that in larger groups of mankind the families exhibiting the particular anomaly had a greater probability of surviving than others.
However this may be, the occurrence of these variations shows that man cannot be assumed to have a stable form. It is of course an entirely open question, how long a time may be required to fix any one of the variations that we are here discussing.
The general tendency of anthropological inquiry has been to assume the permanence of the anatomical characteristics of the present races, beginning with the European races of the early neolithic times. Kollmann, the most pronounced advocate of this theory, claims that the oldest remains of man found in the neolithic deposits of Europe represent types which are still found unchanged among the modern civilized population of the continent. He has tried to identify all the varieties found in the neolithic prehistoric population with those living at the present time.
All studies of the distribution of head-forms and of other anthropometric traits have shown uniformity over considerable continuous areas and through long periods; and the natural inference has been that heredity controls anthropometric forms, and that these are therefore stable (Deniker).
There is only one exception to this rule. In all cases in which the anthropometric traits undergo very considerable changes during the period of growth, the influence of favorable or unfavorable causes makes itself felt. The investigations conducted by Gould and Baxter during the war of the Rebellion have shown that the representatives of European nationalities born in America have statures higher than the representatives of the same nationalities born in Europe; and it has been assumed that better nutrition, or perhaps better hygienic and economic conditions in general, might increase the stature of a people. These conclusions were confirmed by Bowditch’s measurements of the school-children of Boston, and by Peckham’s anthropometric work in Milwaukee. These changes in stature, due to changed conditions, have recently been demonstrated also in Europe, where Ammon has shown that the population of Baden has materially increased in size during the last thirty years. Other corroborative evidence has been obtained from the study of various social classes, in which Bowditch found an increase of stature, beginning with the children of unskilled laborers, and increasing among those of skilled laborers, members of the mercantile class and of the professional class; and from the observations showing a correlation between the character of streets occupied by the well-to-do and the poor, and the stature of their inhabitants (Ripley). Nevertheless these changes of stature were not interpreted as changes in type, because they may well be understood to be due to the elimination of retarding influences, which prevent many individuals from attaining their normal growth.
The results of the observations on stature are substantiated by other anthropometrical studies of various occupations. The best-authenticated fact, because based on the greatest number of observations, is the difference in type between sailors and soldiers who were measured during the war of the Rebellion. It was found that sailors had legs as long as those of the negroes, and correspondingly a shorter trunk, while their arms were equally as long as those of the soldiers of the army. We may also call to mind the investigations carried on in the gymnasiums of our colleges, which show that a series of measurements which depend largely upon the functions of groups of muscles change very rapidly under the influence of practice. It will be acknowledged at once that differences in the use of muscles during childhood, and continued in later life, must result in differences of structure, either permanent, or at least temporary.
A study of the conditions of growth shows how such changes in the form of the body must develop. Setting aside the prenatal development, we find that at the time of birth some parts of the body are so fully developed that they are not far removed from their final size, while others are quite undeveloped. Thus the skull is, comparatively speaking, large at the time of birth, grows rapidly for a short time, but very soon approaches its full size, and then continues to grow very slowly. The limbs, on the other hand, grow rapidly for many years. Other organs do not begin their rapid development until much later in life. Thus it happens that retarding or accelerating influences acting upon the body at different periods of growth may have quite different results. After the head has nearly completed its growth, retarding influences may still influence the length of the limbs. The face, which grows rapidly for a longer period than the cranium, can be influenced later than the latter. In short, the influence of environment may be the more marked, the less developed the organ that is subject to it. Data on the unequal rate of growth of different parts of the body have been furnished by Weissenberg.
The influence of retardation, so far as it has been studied, seems to be lasting. In other words, a retardation in development is never completely made good by long-continued development. When a child, through unfavorable influences, has grown slowly during a number of years, it will probably continue to grow longer than other, normal children; but the total amount of its growth will always remain too small (Boas and Wissler). On the other hand, children whose development has been accelerated will reach the adult stage early, but nevertheless the total amount of their growth will be relatively great. It follows from this consideration of the effect of retardation and of difference in period, that not only the absolute size, but also the relative proportions, of the body, must be influenced by periods of retardation or acceleration.