Fig. 30.—Skull of Phororhacos Compared with that of the Race-horse Lexington.

We can only speculate on the food of these great birds, and for aught we know to the contrary they may have caught fish, fed upon carrion, or used their powerful feet and huge beaks for grubbing roots; but if they were not more or less carnivorous, preying upon such reptiles, mammals and other birds as came within reach, then nature apparently made a mistake in giving them such a formidable equipment of beak and claw. So far as habits go we might be justified in calling them cursorial birds of prey.

Fig. 31.—Leg of a Horse Compared with that of the Giant Moa.

We really know very little about these Patagonian giants, but they are interesting not only from their great size and astounding skulls, but because of the early age (Miocene) at which they lived and because in spite of their bulk they are in nowise related to the ostriches, but belong near the heron family. As usual, we have no idea why they became extinct, but in this instance man is guiltless, for they lived and died long before he made his appearance, and the ever-convenient hypothesis "change of climate" may be responsible for their disappearance.

Something, perhaps, remains to be said concerning the causes which seem to have led to the development of these giant birds, as well as the reasons for their flightless condition and peculiar distribution, for it will be noticed that, with the exception of the African and South American ostriches the great flightless birds as a rule are, and were, confined to uninhabited or sparsely populated islands, and this is equally true of the many small, but equally flightless birds. It is a seemingly harsh law of nature that all living beings shall live in a more or less active struggle with each other and with their surroundings, and that those creatures which possess some slight advantage over their fellows in the matter of speed, or strength, or ability to adapt themselves to surrounding conditions, shall prosper at the expense of the others. In the power of flight, birds have a great safeguard against changes of climate with their accompanying variations in the supply of food, and, to a lesser extent, against their various enemies, including man. This power of flight, acquired early in their geological history, has enabled birds to spread over the length and breadth of the globe as no other group of animals has done, and to thrive under the most varying conditions, and it would seem that if this power were lost it must sooner or later work harm. Now to-day we find no great wingless birds in thickly populated regions, or where beasts of prey abound; the ostriches roam the desert wastes of Arabia, Africa and South America where men are few and savage beasts scarce, and against these is placed a fleetness of foot inherited from ancestors who acquired it before man was. The heavy cassowaries dwell in the thinly inhabited, thickly wooded islands of Malaysia, where again there are no large carnivores and where the dense vegetation is some safeguard against man; the emu comes from the Australian plains, where also there are no four-footed enemies[11] and where his ancestors dwelt in peace before the advent of man. And the same things are true of the Moas, the Æpyornithes, the flightless birds of Patagonia, the recent dodo of Mauritius and the solitaire of Rodriguez, each and all of which flourished in places where there were no men and practically no other enemies. Hence we deduce that absence of enemies is the prime factor in the existence of flightless birds,[12] although presence of food is an essential, while isolation, or restriction to a limited area, plays an important part by keeping together those birds, or that race of birds, whose members show a tendency to disuse their wings. It will be seen that such combinations of circumstances will most naturally be found on islands whose geological history is such that they have had no connection with adjacent continents, or such a very ancient connection that they were not then peopled with beasts of prey, while subsequently their distance from other countries has prevented them from receiving such population by accident in recent times and has also retarded the arrival of man.

[11] The dingo, or native dog, is not forgotten, but, like man, it is a comparatively recent animal.

[12] Note that in Tasmania, which is very near Australia, both in space and in the character of its animals, there are two carnivorous mammals, the Tasmanian "Wolf" and the Tasmanian Devil, and no flightless birds.

Once established, flightlessness and size play into one another's hands; the flightless bird has no limit placed on its size[13] while granted a food supply and immunity from man; the larger the bird the less the necessity for wings to escape from four-footed foes. So long as the climate was favorable and man absent, the big, clumsy bird might thrive, but upon the coming of man, or in the face of any unfavorable change of climate, he would be at a serious disadvantage and hence whenever either of these two factors has been brought to bear against them the feathered giants have vanished.