The robin has been mentioned as a slow migrant, and as a species it takes 78 days to make the 3,000-mile trip from Iowa to Alaska, a stretch of country that is crossed by advancing spring in 68 days. In this case, however, it does not mean that individual robins are necessarily slow, for probably the northward movement of the species depends upon the continual advance of birds from the rear, the first individuals arriving in a suitable locality remaining to nest, while the northward movement of the species is continued by those still to come.
Special interest attaches to the great variation in the speed at which birds travel in different sections of the broad flyway extending from the Gulf of Mexico to the Arctic Ocean, by way of the Mississippi and Mackenzie Valleys. The blackpolled warbler furnishes an excellent example ([fig. 6]). This species winters in north-central South America and migrates in April across the West Indies to Florida. From this point some individuals fly northwest to the Mississippi Valley, north to Manitoba, northwest to the Mackenzie River, and thence almost due west to western Alaska. In tracing the long route of these birds it is found that a fairly uniform average speed of 30 to 35 miles a day is maintained from the Gulf to Minnesota. Then comes a spurt, for a week later the blackpolls have reached the central part of the Mackenzie Valley and by the following week they are observed in northwestern Alaska. During the latter part of the journey, therefore, many individuals must average more than 200 miles a day. They use 30 days in traveling from Florida to southern Minnesota, a distance of about 1,000 miles, and scarcely half that time to cover the remaining 2,500 miles to Alaska. It should be noted that the increased speed is directly associated with the change in direction, the north-and-south course in the Mississippi Valley being accomplished slowly while the northwesterly course across Canada is made at a much greater speed. Increased speed across western Canada to Alaska is also shown by many other birds. A study of all species traveling up the Mississippi Valley indicates an average speed of about 23 miles a day. From southern Minnesota to southern Manitoba 16 species maintain an average speed of about 40 miles a day. From that point to Lake Athabaska, 12 species travel at an average speed of 72 miles a day; while 5 others travel to Great Slave Lake at 116 miles a day; and another 5 species cover 150 miles a day to reach Alaska. This change is in correlation with a corresponding variation in the isothermal lines, which turn northwestward west of the Great Lakes.
Figure 6.—Migration of the blackpolled warbler. The solid isochronal lines show the places at which these birds arrive at the same time. As the birds move northward these lines become farther apart, showing that the warblers move faster with the advance of spring. From April 30 to May 10 the average speed is about 30 miles a day, while from May 25 to May 30 it is increased to more than 200 miles.
As has been previously indicated, the advance of spring in the northern interior is much more rapid than in the Mississippi Valley and on the Gulf coast. In other words, in the North spring comes with a rush and during the height of the migration season in Saskatchewan the temperature in the southern part of the Mackenzie Valley just about equals that in the Lake Superior area, which is 700 miles farther south. Such conditions, coupled with the diagonal course of the birds across this region of fast-moving spring, exert a great influence on migration and are the chief factors in the acceleration of speed of travel.
Variations in speed of migration in different parts of the country are illustrated also by the movements of the cliff swallow ([fig. 3]), which breeds from Mexico to Alaska and winters in Brazil and Argentina. It would be expected in spring to appear in the United States first in Florida and Texas then in the southern Rocky Mountain region, and finally on the Pacific coast. As a matter of fact, however, the earliest spring records come from north-central California, where the bird usually is common before the first arrivals are observed in Texas or Florida. The route taken, for many years a migration problem, was solved when it was found that these swallows went around the Gulf of Mexico rather than across it. The isochronal lines on the map show the more rapid advance along the Pacific coast. By March 20, when the vanguard has not quite reached the lower Rio Grande in Texas, the species is already north of San Francisco in California.
Altitude at which birds travel
At one time students of bird migration held firmly to the theory that normal migration takes place at heights above 15,000 feet, reasoning (somewhat uncertainly) that flying becomes easier as altitude is gained. Since the development of the airplane, however, and with it man's exploration of the upper regions of the air, it has become common knowledge that rarified atmosphere adds greatly to the difficulties of flight. This is due not only to the reduction in oxygen (whether for gasoline engine or the lungs of a bird) but also to the lack of buoyancy of the rarified air. Such birds as vultures, pelicans, cranes, and some of the hawks feel this the least, since compared with body weight the supporting surface of their wings is very great, but for the smaller and shorter-winged birds lack of buoyancy at high altitudes presents a difficult obstacle in flight. Even when flying close to the earth, small birds have to keep their wings in rapid motion.
Another postulate favoring the high-altitude flying theory was that the wonderful vision of birds was their sole guidance during migratory flights; and to keep landmarks in view the birds were obliged to fly high, particularly when crossing wide areas of water. This will be considered in greater detail under Orientation (p. 28), so here it will be sufficient to say that birds rely only in part upon vision to guide them on migration. Also, it is to be remembered that there are definite physical limitations to the range of visibility even under perfect atmospheric conditions. Chief of these is the curvature of the earth's surface. Thus, if birds crossing the Gulf of Mexico to Louisiana and Florida flew at a height of 5 miles, they would still be unable to see a third of the way across. And yet this trip is made twice each year by thousands of thrushes, warblers, and others.
Actual knowledge of the altitude of migratory flight is scanty, though estimates obtained by means of the telescope, and still more accurate data resulting from altimeter observation from airplanes, are slowly accumulating. It is, of course, obvious that some birds that cross mountain ranges during migration must attain a great altitude. Observers at an altitude of 14,000 feet in the Himalayas have recorded storks and cranes flying so high that they could be seen only through field glasses. Being beyond the range of unaided vision they must have been at least 6,000 feet above the observers, or at an actual altitude of 20,000 feet above sea level. Such cases, however, are exceptional as aviators have reported that they rarely meet birds above an altitude of 5,000 feet.