The movements of the western tanager show a migration route that is in some ways remarkable. The species breeds in the mountains from the northern part of Baja California and western Texas north to northeastern British Columbia and southwestern Mackenzie. Its winter range is in two discontinuous areas—southern Baja California and eastern Mexico south to Guatemala ([fig. 21]). On the spring migration the birds enter the United States about April 20, appearing first in western Texas and the southern parts of New Mexico and Arizona ([fig. 22]). By April 30 the vanguard has advanced evenly to an approximate east-and-west line across central New Mexico, Arizona, and southern California. But by May 10 the easternmost birds have advanced only to southern Colorado, while those in the far west have reached northern Washington. Ten days later the northward advance of the species is shown as a great curve, extending northeastward from Vancouver Island to central Albert and thence southeastward to northern Colorado. Since these tanagers do not reach northern Colorado until May 20, it is evident that those present in Alberta on that date, instead of traveling northward through the Rocky Mountains, which from the location of their summer and winter homes would seem to be the natural route, reached there by the Pacific coast route to southern British Columbia and thence across the mountains, despite the fact that these are still partly covered with snow at that time.
Pacific oceanic route
The route of the Pacific golden plover is fully as interesting and as remarkable as the elliptical course followed by its eastern cousin ([fig. 18]). The breeding range of the eastern golden plover extends through Arctic America west to the northern coast of Alaska where, in the vicinity of Point Barrow, it meets the nesting grounds of the Pacific form, which is really an Asiatic subspecies. It breeds chiefly in the Arctic coast region of Siberia and merely overflows onto the Alaskan coast, some of the birds probably migrating south along the coast of Asia to winter quarters in Japan, China, India, Australia, New Zealand, and Oceania, including the Hawaiian Islands, the Marquesas Islands, and the Low Archipelago. Golden plovers in migration have been observed at sea on a line that apparently extends from these islands to the Aleutians, and it therefore appears certain that at least some of the Alaskan birds make a nonstop flight across a landless sea from Alaska to Hawaii. While it would seem incredible that any birds could lay a course so straight as to attain these small oceanic islands, 2,000 miles south of the Aleutians, 2,000 miles west of Baja California, and nearly 4,000 miles east of Japan, the evidence admits only the conclusion that year after year this transoceanic round-trip journey between Alaska and Hawaii is made by considerable numbers of golden plovers.
Figure 22.—Migration of the western tanager. The birds that arrive in eastern Alberta by May 20 do not travel northward along the eastern base of the Rocky Mountains, as in that region the van has then only reached northern Colorado. Instead the isochronal lines indicate that they migrate north through California, Oregon, and Washington, and then cross the mountains of British Columbia.
The Pacific oceanic route probably is used also by the arctic terns that breed in Alaska, and possibly by those from the more western tern colonies of Canada. This species is of regular occurrence on the western coasts of both the United States and South America, indicating that the western representatives travel southward to the Antarctic winter quarters without the spectacular migration features that appear to characterize the flight of those from the eastern part of the continent ([fig. 8]).
Arctic routes
In the discussion of the migration of the Arctic tern (p. 38) it was noted that this species makes a very distinct west-to-east movement across northern Canada, continuing the flight eastward across the Atlantic Ocean toward the western coast of Europe. It seems likely that there are other species, including the parasitic jaeger, that regularly breed in the northern part of the Western Hemisphere but migrate back to the Old World for their winter sojourn. Some others, as the red-legged kittiwake and Ross's gull, remain near the Arctic region throughout the year, retreating southward in winter only a few hundred miles. The emperor goose in winter is found only a relatively short distance south of its breeding grounds, and eider ducks, although wintering in latitudes well south of their nesting areas, nevertheless remain farther north than do the majority of other species of ducks.
The routes followed by these birds are chiefly coastwise, and in the final analysis may be considered as being tributary either to the Atlantic or to the Pacific coast routes. The passage of gulls, ducks, the black brant, and other water birds at Point Barrow, Alaska, and at other points on the Arctic coast, has been noted by several observers, and from present knowledge it may be said that the best defined Arctic route in North America is the one that follows around the coast of Alaska.
