Various forms of Foraminifera.
a. Lagena striata. a′. Nodosaria rugosa. b. Marginulina raphanus.
b′. Longitudinal section of shell of ditto. c. Polystomella crispa, with its pseudopodia protruded.
d. Nummulites lenticularis, shown in horizontal section. e. Cassidulina lævigata.
f. Textularia globulosa. g. Miliolina seminulum. g′. Animal of Miliolina removed from its shell.
The amazing variety of form of the Foraminifera is no less remarkable than the elegance of their delicately chiselled shells, and may well be called immense, as no less than 2,400 living and fossil species have already been distinguished by naturalists, and a far greater number is probably still nameless and unknown. Though generally so minute that the diameter of the pores through which they protrude their filaments usually only ranges from 1/3000 to 1/10000 of an inch yet the diminutive world of the Foraminifera has also its giants, particularly among the fossil species, such as the Nummulites, which occur in such prodigious numbers in the limestone of the Egyptian pyramids, and whose flattened lenticular coin-like forms (d) attain the comparatively gigantic diameter of several inches. Thus the material with which the proud Pharaohs of the Nile constructed their colossal tombs was originally piled up at the bottom of the sea by countless generations of shell-cased Protozoa.
The Foraminifera are among the oldest inhabitants of our globe,[T] and as the present ocean contains them in countless multitudes, thus have they swarmed in the waters of the primeval seas from the first dawn of creation, and piled up the monuments of their existence in vast strata of limestone. A great part of the rocky belt from Rügen to the Danish isles, the white chalk cliffs which, beginning in England, extend through France as far as Southern Spain, are chiefly composed of the shells of Foraminifera, and the zone of Nummulite limestone, which served to build the huge quadrilateral monument of Cheops, forms a band, often 1,800 miles in breadth, and frequently of enormous thickness, from the Atlantic shores of Europe and Africa through Western Asia up to North India and China; enough to satisfy the most extravagant architectural folly of millions of despots. So important is the part which these beings, individually so minute, have performed and still perform in the geological annals of the globe.
[T] The Eozoon canadense, the oldest of known organic remains, found in the Upper Laurentian series, which preceded the Cambrian formation, is a Foraminifer. Millions of years must have passed since it first felt and moved.
Many of these "minims of nature" consist of only one chamber, and hence are called unilocular or monothalamous; but a vast proportion consist of several chambers, and hence are called multilocular or polythalamous. The latter, however numerous their chambers or seemingly complex their structure, always originate as a single shell. The primitive jelly-sphere, or first sarcode segment, secretes around itself its appropriate calcareous envelope. Having grown too large for its habitation, it protrudes a portion of itself without, and thus forms a second segment. If by a process of spontaneous fission this segment becomes quite detached from its parent, and repeats the life and method of reproduction of the latter, a series of monothalamous shells will be formed. But if by means of a sarcode band the primitive segment maintains its connection with its immediate offspring, and this, repeating the reproductive process, does the same, a compound shell will, of course, be the result.
Among the microscopic denizens of the ocean, the Polycystina rival the Foraminifera both by their number and their wonderful elegance of form and structure. Their body consists of the same viscid homogeneous plastic mass, termed "sarcode" by the naturalists; like them they are capable of protruding it through the foramina with which their shell is pierced, and consequently they are ranked with them among the Rhizopods, or root-footed animalcules, that form the lowest order of the Protozoa, the lowest class of the animal world.
Polycystina.
a. Podocyrtis Schomburgkii. b. Haliomma Humboldtii.