Fig. 326.—Tarpon, Megalops atlanticus, much reduced. (After Goode.)

Fam. 6. Albulidae.—Margin of the upper jaw formed by the praemaxillaries and the maxillaries, the latter movably articulated above the former to the ethmoid. Parietal bones separating the supraoccipital from the frontals; suboperculum large; interoperculum small and entirely or nearly entirely hidden below the praeoperculum. Basis cranii double. Praemaxillaries, mandible, vomer, palatines, pterygoid, parasphenoid, glossohyal, and pharyngeals toothed. Ribs sessile, inserted behind and somewhat below small parapophyses, which are absent or merely indicated on the anterior vertebrae, and gradually increase in size towards the caudal region; these parapophyses, as well as the neural and haemal arches, are autogenous bones; epineurals, no epipleurals. Pectorals low down, folding like the ventrals. Post-temporal trifid, the upper branch attached to the epiotic, the median to the squamosal, the lower to the opisthotic; post-clavicle large (formed of three pieces); scapular foramen between scapula and clavicle; pterygials well developed, two in contact with coracoid. Ventrals with 10 to 14 rays. Branchiostegal rays 6 to 14. Air-bladder large, not communicating with the ear.

Elongate fusiform Fishes, covered with large silvery scales forming regular longitudinal series; head naked; mouth rather small, with thick lips; gill-openings wide. Dorsal fin originating in front of ventrals; anal short; caudal well developed, forked.

Fig. 327.—Albula conorhynchus. ¼ nat. size.

The type of this family, Albula or Butirinus, is remarkable among Teleosts in possessing a rudiment of a conus arteriosus to the heart, provided with two rows of valvules.[[642]] Its single species inhabits all the warm and tropical seas. Prof. Gilbert has ascertained that the young pass through a metamorphosis, analogous to that of the Eels. The "Leptocephalid" described as Esunculus by Kaup is probably a larval Albula.

Fig. 328.—Larva of Albula conorhynchus. (After Gilbert.)

The deep-sea Japanese Pterothrissus (Bathythrissa) must be placed in this family; its skeleton is very similar to that of Albula. It differs in the elongate dorsal fin, in the presence of small teeth on the maxillary bone, and in the small number of branchiostegal rays (6 instead of 12 to 14).