Fam. 3. Lipogenyidae.—Similar to the preceding in shape and in the position of the dorsal fin, but with a toothless, roundish, inferior, suctorial mouth, and with the short dorsal and the long anal formed partly of spines and partly of soft rays. Head and body covered with minute scales; lateral line nearer the dorsal than the ventral profile. Ventrals with 3 spines and 7 soft rays. A single species, Lipogenys gillii, from the North Atlantic, 865 fathoms.
Fig. 380.—Lipogenys gillii. (After Goode and Bean.)
Fam. 4. Notacanthidae.—Body elongate, covered with very small cycloid scales, the tail tapering to a point, without caudal fin; head scaly; mouth small, inferior, bordered by the praemaxillaries only; jaws toothed; no suborbitals; praeoperculum small; post-temporal replaced by ligament. Dorsal fin formed of a series of short disconnected spines; anal very long, formed partly of spines and partly of soft rays, extending to the end of the tail. Ventrals with 1 to 5 spines and 7 to 10 soft rays.
Two genera: Notacanthus, with the ventrals connate or confluent and with 6 to 12 dorsal spines; and Polyacanthonotus, with the ventrals separated and 27 to 38 dorsal spines. Nine species, from the Mediterranean, the Atlantic, and the Pacific, at depths of 400 to 1875 fathoms.
Fig. 381.—Notacanthus bonapartii, ½ nat. size. (After Vaillant.)
Fam. 5. Fierasferidae.—Body elongate or extremely attenuate, naked, the tail tapering to a point or truncate, without distinct caudal fin; mouth small, inferior, bordered by the praemaxillaries; jaws toothed; no suborbitals; praeoperculum well developed. Dorsal and anal fins very long, extending to the end of the tail, and formed entirely of soft rays. Ventral fins absent. Vent situated immediately behind the gill-opening. Air-bladder with a muscular apparatus for dilatation of its anterior part.
A single genus, Fierasfer, with about 10 species, distributed over nearly all the warm and tropical seas, rarely found as far north as the west coast of Ireland. Encheliophis, without pectoral fins, is the larval form of Fierasfer.
Fierasfer spends the greater part of its existence in the interior of Holothurians and other Echinoderms as well as in bivalve Mollusca. It has been observed to enter Holothurians by the posterior or anal aperture, either head first or tail foremost, in the latter case availing itself of the suction which takes place alternately with the expulsion of water by that orifice; it remains near the anus, from which it projects its head in search of food outside its host. It is neither a true parasite nor a commensal or mutualist, in the sense given to these terms by Van Beneden, but simply a lodger, "inquilino," as Emery puts it. Semper, however, regards Encheliophis vermicularis as a true parasite, feeding on the viscera of the Holothurian in which it lives. Putnam has examined eight specimens of a Fierasfer from the Bay of Panama, which were obtained alive from pearl oysters, and also one beautifully enclosed in a pearly covering deposited upon it by the oyster; a similar specimen is preserved in the British Museum.