The tubular fangs of the Viperidæ are inserted on the posterior extremity of the much abbreviated and erectile maxillary bone, which bears no other teeth. The Proteroglyphs (Cobras, Coral-snakes, Sea-snakes) and the Solenoglyphs (Vipers, Pit-vipers, Rattlesnakes) may be regarded as the diverging extremes in the development of the poison apparatus, both culminating in forms with tubular fangs, the former as derived directly from the Aglyphs (harmless snakes), the latter from the Opisthoglyphs, likewise evolved out of the Aglyphs. That the insertion of the poison fangs of the Viperidæ is really on the posterior extremity of the maxillary bone is evident from the condition of the bone in its recumbent position, especially in the African Viper, Causus, which in several respects departs less markedly from the Colubrid type than our European Vipers.
The poison fangs of the Viperidæ appear to be movable, folding in the mouth when at rest, and erected, or even thrust forward, when ready to act. This, however, is simply due to the mobility of the maxillary bone, to which they are ankylosed as in all other snakes. There are normally two equally-developed fangs, close together and side by side, to each maxillary, followed by several replacement fangs loosely attached behind them, usually in two series of four. When the two fangs are in situ, they of course both function in the act of biting, although only one is in relation with the single poison duct; often, however, there is only one fang in position, either the right or the left, the place of the other being indicated by a shallow socket which will soon be filled by one of the posterior reserve fangs moving forward and becoming ankylosed to the bone. Snake-charmers who extract the poison fangs of the snakes they use for their performances have therefore to renew the operation frequently, unless they amputate the bone on which the fangs are inserted, an injury which the creature does not long survive.
The dentition of the snakes in which the maxillary bone is not movable vertically falls under three divisions: the Aglyphs, in which the teeth are all solid; the Opisthoglyphs, in which one or more (usually two) of the hindermost teeth are provided with a groove; and the Proteroglyphs, in which grooved or canaliculated teeth are situated in front, followed or not by solid teeth. Beyond these three principal divisions, the dentition furnishes important characters for the classification, although that importance has sometimes been over-estimated. The maxillary teeth may be equal in length (Isodonts), or the anterior the longer (Lycodonts), or the posterior the longer, increasing gradually in size (Coryphodonts) or abruptly, without (Syncranterians) or with a diastema, or break, in front of them (Diacranterians). These categories are, however, so completely connected as to preclude their use in taxonomy beyond helping to define genera. The number of maxillary teeth and the relative proportions and disposition of the mandibular teeth also afford useful generic characters.
The European genera may be arranged as follows, according to the dentition:
I. Teeth few, disposed in a transverse series in the upper jaw only
Typhlops.
II. Teeth in both jaws and on the palatines and pterygoids.
A. A series of solid teeth along the maxillary; no grooved teeth.
1. Anterior maxillary and mandibular teeth longest; 9 or 10 maxillary teeth
Eryx.