In the first place, besides the works of Darwin himself, there are a number of others which have already and very admirably expounded the evidences, both of organic evolution as a fact, and of natural selection as a cause. Therefore, in the present treatise it seemed needless to go beyond the ground which was covered by my original lectures, namely, a condensed and connected, while at the same time a critical statement of the main evidences, and the main objections, which have thus far been published with reference to the distinctively Darwinian theory. Indeed while re-casting this portion of my lectures for the present publication, I have felt that criticism might be more justly urged from the side of impatience at a reiteration of facts and arguments already so well known. But while endeavouring, as much as possible, to avoid overlapping the previous expositions, I have not carried this attempt to the extent of damaging my own, by omitting any of the more important heads of evidence; and I have sought to invest the latter with some measure of novelty by making good what appears to me a deficiency which has hitherto obtained in the matter of pictorial illustration. In particular, there will be found a tolerably extensive series of woodcuts, serving to represent the more important products of artificial selection. These, like all the other original illustrations, have been drawn either direct from nature or from a comparative study of the best authorities. Nevertheless, I desire it to be understood that the first part of this treatise is intended to retain its original character, as a merely educational exposition of Darwinian teaching—an exposition, therefore, which, in its present form, may be regarded as a compendium, or hand-book, adapted to the requirements of a general reader, or biological student as distinguished from those of a professed naturalist.

The case, however, is different with the second instalment, which will be published at no very distant date. Here I have not followed with nearly so much closeness the material of my original lectures. On the contrary, I have had in view a special class of readers; and, although I have tried not altogether to sacrifice the more general class, I shall desire it to be understood that I am there appealing to naturalists who are specialists in Darwinism. One must say advisedly, naturalists who are specialists in Darwinism, because, while the literature of Darwinism has become a department of science in itself, there are nowadays many naturalists who, without having paid any close attention to the subject, deem themselves entitled to hold authoritative opinions with regard to it. These men may have done admirable work in other departments of natural history, and yet their opinions on such matters as we shall hereafter have to consider may be destitute of value. As there is no necessary relation between erudition in one department of science and soundness of judgment in another, the mere fact that a man is distinguished as a botanist or zoologist does not in itself qualify him as a critic where specially Darwinian questions are concerned. Thus it happens now, as it happened thirty years ago, that highly distinguished botanists and zoologists prove themselves incapable as judges of general reasoning. It was Darwin’s complaint that for many years nearly all his scientific critics either could not, or would not, understand what he had written—and this even as regarded the fundamental principles of his theory, which with the utmost clearness he had over and over again repeated. Now the only difference between such naturalists and their successors of the present day is, that the latter have grown up in a Darwinian environment, and so, as already remarked, have more or less thoughtlessly adopted some form of Darwinian creed. But this scientific creed is not a whit less dogmatic and intolerant than was the more theological one which it has supplanted; and while it usually incorporates the main elements of Darwin’s teaching, it still more usually comprises gross perversions of their consequences. All this I shall have occasion more fully to show in subsequent parts of the present work; and allusion is made to the matter here merely for the sake of observing that in future I shall not pay attention to unsupported expressions of opinion from any quarter: I shall consider only such as are accompanied with some statement of the grounds upon which the opinion is held. And, even as thus limited, I do not think it will be found that the following exposition devotes any disproportional amount of attention to the contemporary movements of Darwinian thought, seeing, as we shall see, how active scientific speculation has been in the field of Darwinism since the death of Mr. Darwin.


Leaving, then, these post-Darwinian questions to be dealt with subsequently, I shall now begin a systematic résumé of the evidences in favour of the Darwinian theory, as this was left to the world by Darwin himself.

There is a great distinction to be drawn between the fact of evolution and the manner of it, or between the evidence of evolution as having taken place somehow, and the evidence of the causes which have been concerned in the process. This most important distinction is frequently disregarded by popular writers on Darwinism; and, therefore, in order to mark it as strongly as possible, I will effect a complete separation between the evidence which we have of evolution as a fact, and the evidence which we have as to its method. In other words, not until I shall have fully considered the evidence of organic evolution as a process which somehow or another has taken place, will I proceed to consider how it has taken place, or the causes which Darwin and others have suggested as having probably been concerned in this process.

Confining, then, our attention in the first instance to a proof of evolution considered as a fact, without any reference at all to its method, let us begin by considering the antecedent standing of the matter.


First of all we must clearly recognise that there are only two hypotheses in the field whereby it is possible so much as to suggest an explanation of the origin of species. Either all the species of plants and animals must have been supernaturally created, or else they must have been naturally evolved. There is no third hypothesis possible; for no one can rationally suggest that species have been eternal.

Next, be it observed, that the theory of a continuous transmutation of species is not logically bound to furnish a full explanation of all the natural causes which it may suppose to have been at work. The radical distinction between the two theories consists in the one assuming an immediate action of some supernatural or inscrutable cause, while the other assumes the immediate action of natural—and therefore of possibly discoverable—causes. But in order to sustain this latter assumption, the theory of descent is under no logical necessity to furnish a full proof of all the natural causes which may have been concerned in working out the observed results. We do not know the natural causes of many diseases; but yet no one nowadays thinks of reverting to any hypothesis of a supernatural cause, in order to explain the occurrence of any disease the natural causation of which is obscure. The science of medicine being in so many cases able to explain the occurrence of disease by its hypothesis of natural causes, medical men now feel that they are entitled to assume, on the basis of a wide analogy, and therefore on the basis of a strong antecedent presumption, that all diseases are due to natural causes, whether or not in particular cases such causes happen to have been discovered. And from this position it follows that medical men are not logically bound to entertain any supernatural theory of an obscure disease, merely because as yet they have failed to find a natural theory. And so it is with biologists and their theory of descent. Even if it be fully proved to them that the causes which they have hitherto discovered, or suggested, are inadequate to account for all the facts of organic nature, this would in no wise logically compel them to vacate their theory of evolution, in favour of the theory of creation. All that it would so compel them to do would be to search with yet greater diligence for the natural causes still undiscovered, but in the existence of which they are, by their independent evidence in favour of the theory, bound to believe.

In short, the issue is not between the theory of a supernatural cause and the theory of any one particular natural cause, or set of causes—such as natural selection, use, disuse, and so forth. The issue thus far—or where only the fact of evolution is concerned—is between the theory of a supernatural cause as operating immediately in numberless acts of special creation, and the theory of natural causes as a whole, whether these happen, or do not happen, to have been hitherto discovered.