Movements of the metacarpal bones of the Fingers upon the Wrist.

The movements of the thumb, through the medium of its metacarpal bone, upon the wrist are much more free than those of any of the other fingers. The middle finger, indeed, has its metacarpal bone set upon the wrist so fixedly as to admit of scarcely any such movement. The forefinger can be thus moved a little; the ring finger more and the little finger still more.

You may easily prove this by taking the knuckles or heads of the respective metacarpal bones of one hand between the fingers and thumb of the other hand, when you will find that you can press the knuckle of the little finger backwards and forwards in a very perceptible manner. Then try the knuckle of the ring finger; the movement is distinct, though not so free as in the case of the little finger. The knuckle of the forefinger you will find to be almost fixed; and in that of the middle finger you will be unable to perceive any movement at all.

In fact the joints of the metacarpal bones of the fingers with the wrist resemble those of the outer four toes with the tarsus; and the drawings of these joints of the foot (figs. [22] to 25) will serve sufficiently well to illustrate those of the hand.

These movements of the metacarpal, or knuckle, bones upon the wrist enable us to increase or diminish the hollow of the palm by bringing its edges more or less forward. Thus, when we make a cup of the hand we bring forward the metacarpal bones of the thumb and the little finger, wrinkling the skin of the palm; and when we spread the hand open we carry those bones backwards, rendering the skin of the palm tense.

These movements, moreover, enable us to bring the little fingers and the thumb more easily into contact.

Have you ever wondered what advantage is gained by the fingers and thumb all differing from one another in length; or don’t you take the trouble to reflect upon little matters of this sort? If you have, I would ask you now to remark that there is, in the several fingers, a relation between their shortness, their position near the edge of the hand, and the amount of mobility of their metacarpal bones upon the wrist. Thus the finger which is in the middle of the hand is the longest, and its metacarpal is the most fixed. The fore-finger is not quite so long; and its metacarpal is rather less immovable. The ring-finger comes next in shortness and in the mobility of its metacarpal. Then the little finger; and the thumb which is much shorter than any other has also its metacarpal much more moveable.

Fig. [68].

Observe, further, that, when the fingers and thumb are separated from one another, and then bent, the middle knuckle-bone remains stationary, but the others are advanced a little forwards, each to an extent proportionate to its mobility upon the wrist, and to the shortness of the finger. The fore-finger is, by this means, advanced a little, the ring and the little fingers more, and the thumb most of all. And the result is, that the tips of the fingers and the thumb come all to a level, and form, with the palm, a great hollow in which we can grasp any substance, a cricket-ball, for instance, and hold it very firmly. The length of the several fingers and the thumb is, therefore, just so regulated, in relation to their mobility upon the wrist, as to give us this power.