[167] "The best state of weather for a good run," says my correspondent, "is clear days, thawing fast in the daytime and freezing well at night, with a gentle west or northwest wind; though we sometimes have clear, fine, thawing days followed by frosty nights, without a good run of sap, I have thought it probable that the irregular flow of sap on different days in the same season is connected with the variation in atmospheric pressure; for the atmospheric conditions above mentioned as those most favorable to a free flow of sap are also those in which the barometer usually indicates pressure considerably above the mean. With a south or southeast wind, and in lowering weather, which causes a fall in the barometer, the flow generally ceases, though the sap sometimes runs till after the beginning of the storm. With a gentle wind, south of west, maples sometimes run all night. When this occurs, it is oftenest shortly before a storm. Last spring, the sap of a sugar orchard in a neighboring town flowed the greater part of the time for two days and two nights successively, and did not cease till after the commencement of a rain storm."
The cessation of the flow of sap at night is perhaps in part to be ascribed to the nocturnal frost, which checks the melting of the snow, of course diminishing the supply of moisture in the ground, and sometimes congeals the strata from which the rootlets suck in water. From the facts already mentioned, however, and from other well-known circumstances—such, for example, as the more liberal flow of sap from incisions on the south side of the trunk—it is evident that the withdrawal of the stimulating influences of the sun's light and heat is the principal cause of the suspension of the circulation in the night.
[168] "The flow ceases altogether soon after the buds begin to swell."—Letter before quoted.
[169] We might obtain a contribution to an approximate estimate of the quantity of moisture abstracted by forest vegetation from the earth and the air, by ascertaining, as nearly as possible, the quantity of wood on a given area, the proportion of assimilable matter contained in the fluids of the tree at different seasons of the year, the ages of the trees respectively, and the quantity of leaf and seed annually shed by them. The results would, indeed, be very vague, but they might serve to check or confirm estimates arrived at by other processes. The following facts are items too loose perhaps to be employed as elements in such a computation.
Dr. Williams, who wrote when the woods of Northern New England were generally in their primitive condition, states the number of trees growing on an acre at from one hundred and fifty to six hundred and fifty, according to their size and the quality of the soil; the quantity of wood, at from fifty to two hundred cords, or from 238 to 952 cubic yards, but adds that on land covered with pines, the quantity of wood would be much greater. Whether he means to give the entire solid contents of the tree, or, as is usual in ordinary estimates in New England, the marketable wood only, the trunks and larger branches, does not appear. Next to the pine, the maple would probably yield a larger amount to a given area than any of the other trees mentioned by Dr. Williams, but mixed wood, in general, measures most. In a good deal of observation on this subject, the largest quantity of marketable wood I have ever known cut on an acre of virgin forest was one hundred and four cords, or 493 cubic yards, and half that amount is considered a very fair yield. The smaller trees, branches, and twigs would not increase the quantity more than twenty-five per cent., and if we add as much more for the roots, we should have a total of about 750 cubic yards. I think Dr. Williams's estimate too large, though it would fall much below the product of the great trees of the Mississippi Valley, of Oregon, and of California. It should be observed that these measurements are those of the wood as it lies when 'corded' or piled up for market, and exceed the real solid contents by not less than fifteen per cent.
"In a soil of medium quality," says Clavé, quoting the estimates of Pfeil, for the climate of Prussia, "the volume of a hectare of pines twenty years old, would exceed 80 cubic mètres [42½ cubic yards to the acre]; it would amount to but 24 in a meagre soil. This tree attains its maximum of mean growth at the age of seventy-five years. At that age, in the sandy earth of Prussia, it produces annually about 5 cubic mètres, with a total volume of 311 cubic mètres per hectare [166 cubic yards per acre]. After this age the volume increases, but the mean rate of growth diminishes. At eighty years, for instance, the volume is 335 cubic mètres, the annual production 4 only. The beech reaches its maximum of annual growth at one hundred and twenty years. It then has a total volume of 633 cubic mètres to the hectare [335 cubic yards to the acre], and produces 5 cubic mètres per year."—Clavé, Études, p. 151.
These measures, I believe, include the entire ligneous product of the tree, exclusive of the roots, and express the actual solid contents. The specific gravity of maple wood is stated to be 75. Maple sap yields sugar at the rate of about one pound wet sugar to three gallons of sap, and wet sugar is to dry sugar in about the proportion of nineteen to sixteen. Besides the sugar, there is a small residuum of "sand," composed of phosphate of lime, with a little silex, and it is certain that by the ordinary hasty process of manufacture, a good deal of sugar is lost; for the drops, condensed from the vapor of the boilers on the rafters of the rude sheds where the sap is boiled, have a decidedly sweet taste.
[170] "The elaborated sap, passing out of the leaves, is received into the inner bark, * * * and a part of what descends finds its way even to the ends of the roots, and is all along diffused laterally into the stem, where it meets and mingles with the ascending crude sap or raw material. So there is no separate circulation of the two kinds of sap; and no crude sap exists separately in any part of the plant. Even in the root, where it enters, this mingles at once with some elaborated sap already there."—Gray, How Plants Grow, § 273.
[171] Ward's tight glazed cases for raising, and especially for transporting plants, go far to prove that water only circulates through vegetables, and is again and again absorbed and transpired by organs appropriated to these functions. Seeds, growing grasses, shrubs, or trees planted in proper earth, moderately watered and covered with a glass bell or close frame of glass, live for months and even years, with only the original store of air and water. In one of Ward's early experiments, a spire of grass and a fern, which sprang up in a corked bottle containing a little moist earth introduced as a bed for a snail, lived and flourished for eighteen years without a new supply of either fluid. In these boxes the plants grow till the enclosed air is exhausted of the gaseous constituents of vegetation, and till the water has yielded up the assimilable matter it held in solution, and dissolved and supplied to the roots the nutriment contained in the earth in which they are planted. After this, they continue for a long time in a state of vegetable sleep, but if fresh air and water be introduced into the cases, or the plants be transplanted into open ground, they rouse themselves to renewed life, and grow vigorously, without appearing to have suffered from their long imprisonment. The water transpired by the leaves is partly absorbed by the earth directly from the air, partly condensed on the glass, along which it trickles down to the earth, enters the roots again, and thus continually repeats the circuit. See Aus der Natur, 21, B. S. 537.
[172] Wilhelm, Der Boden und das Wasser, p. 18. It is not ascertained in what proportions the dew is evaporated, and in what it is absorbed by the earth, in actual nature, but there can be no doubt that the amount of water taken up by the ground, both from vapor suspended in the air and from dew, is large. The annual fall of dew in England is estimated at five inches, but this quantity is much exceeded in many countries with a clearer sky. "In many of our Algerian campaigns," says Babinet, "when it was wished to punish the brigandage of the unsubdued tribes, it was impossible to set their grain fields on fire until a late hour of the day; for the plants were so wet with the night dew that it was necessary to wait until the sun had dried them."—Études et Lectures, ii, p. 212.