Experiments by Meguscher, in Lombardy, led that observer to conclude "that the wood of a living tree maintains a temperature of + 12 degrees or 18 degrees Cent. [= 54 degrees, 56 degrees Fahr.] when the temperature of the air stands at 3 degrees, 7 degrees, and 8 degrees [= 37 degrees, 46 degrees, 47 degrees F.] above zero, and that the internal warmth of the tree does not rise and fall in proportion to that of the atmosphere. So long as the latter is below 18 degrees [= 67 degrees Fahr.], that of the tree is always the highest; but if the temperature of the air rises to 18 degrees, that of the vegetable growth is the lowest. Since then, trees maintain at all seasons a constant mean temperature of 12 degrees [= 54 degrees Fahr.], it is easy to see why the air in contact with the forest must be warmer in winter, cooler in summer than in situations where it is deprived of that influence." [Footnote: Memoria Sur Boschi Della Lombardia, p. 45. The results of recent experiments by Becquerel do not accord with those obtained by Meguscher, and the former eminent physicist holds that "a tree is warmed in the air like any inert body." At the same time he asserts, as a fact well ascertained by experiment, that "vegetables possess in themselves the power or resisting extreme cold for a certain length of time,…. and hence it is believed that there may exist in the organism of plants a force, independent of the conduction of caloric, which resists a degree of cold above the freezing-point." In a following page he cites observations made by Bugeaud, under the parallel of 58 degrees N. L., between the months of November and June, during most of which time, of course, vegetable life was in its deepest lethargy. Bugeaud found that when the temperature of the air was at -34.60 degrees, that of a poplar was only at -29.70 degrees, which certainly confirms the doctrine that trees exercise a certain internal resistance against cold.]

Professor Henry says: "As a general deduction from chemical and mechanical principles, we think no change of temperature is ever produced where the actions belonging to one or both of these principles are not present. Hence, in midwinter, when all vegetable functions are dormant, we do not believe that any heat is developed by a tree, or that its interior differs in temperature from its exterior further than it is protected from the external air. The experiments which have been made on this point, we think, have been directed by a false analogy. During the active circulation of the sap and the production of new tissue, variations of temperature belonging exclusively to the plant may be observed; but it is inconsistent with general principles that heat should be generated where no change is taking place." [Footnote: United States Patent Office Report for 1857, p. 504.]

There can be no doubt that moisture is given, out by trees and evaporated in extremely cold winter weather, and unless new fluid were supplied from the roots by the exercise of some vital function, the tree would be exhausted of its juices before winter was over. But this is not observed to be the fact, and, though the point is disputed, respectable authorities declare that "wood felled in the depth of winter is the heaviest and fullest of sap." [Footnote: Rossmassler, Der Wald, p. 158.] Warm weather in winter, of too short continuance to affect the temperature of the ground sensibly, stimulates a free flow of sap in the maple. Thus, in the last week of December, 1862, and the first week of January, 1863, sugar was made from that tree in various parts of New England. "A single branch of a tree, admitted into a warm room in winter through an aperture in a window, opened its buds and developed its leaves, while the rest of the tree in the external air remained in its winter sleep." [Footnote: Ibid., p. 160.] Like facts are matter of every-day observation in graperies where the vine is often planted outside the wall, the stem passing through an aperture into the warm interior. The roots, of course, stand in ground of the ordinary winter temperature, but vegetation is developed in the branches at the pleasure of the gardener. The roots of forest trees in temperate climates remain, for the most part, in a moist soil, of a temperature not much below the annual mean, through the whole winter; and we cannot account for the uninterrupted moisture of the tree, unless we suppose that the roots furnish a constant supply of water. Atkinson describes a ravine in a valley in Siberia, which was filled with ice to the depth of twenty-five feet. Poplars were growing in this ice, which was thawed to the distance of some inches from the stem. But the surface of the soil beneath it must have remained still frozen, for the holes around the trees were full of water resulting from its melting, and this would have escaped below if the ground had been thawed. In this case, although the roots had not thawed the thick covering of earth above them, the trunks must have melted the ice in contact with them. The trees, when observed by Atkinson, were in full leaf, but it does not appear at what period the ice around their stems had melted.

From these facts, and others of the like sort, it would seem that "all vegetable functions are" not absolutely "dormant in winter, and, therefore, that trees may give out SOME heat even at that season." [Footnote: All evergreens, even the broad-leaved trees, resist frosts of extraordinary severity better than the deciduous trees of the same climates. Is not this because the vital processes of trees of persistent foliage are less interrupted during winter than those of trees which annually shed their leaves, and that therefore more organic heat is developed?

In crossing Mont Cenis in October, 1869, when the leaves of the larches on the northern slope and near the top of the mountain were entirely dead and turned brown, I observed that these trees were completely white with hoar-frost. It was a wonderful sight to see how every leaf was covered with a delicate deposit of frozen aqueous vapor, which gave the effect of the most brilliant silver. On the other band, the evergreen coniferae, which were growing among the larches, and therefore in the same conditions of exposure, were almost entirely free from frost. The contrast between the verdure of the leaves of the evergreens and the crystalline splendor of those of the larches was strikingly beautiful. Was this fact due to a difference in the color and structure of the leaves, or rather is it a proof of a vital force of resistance to cold in the living foliage of the evergreen tree The low temperature of air and soil at which, in the frigid zone, as well as in warmer latitudes under special circumstances, the processes of vegetation go on, seems to necessitate the supposition that all the manifestations of vegetable life are attended with an evolution of heat. In the United States it is common to protect ice, in ice-houses, by a covering of straw, which naturally sometimes contains kernels of grain. These often sprout, and even throw out roots and leaves to a considerable length, in a temperature very little above the freezing-point. Three or four years since I saw a lump of very clear and apparently solid ice, about eight inches long by six thick, on which a kernel of grain had sprouted in an ice-house, and sent half a dozen or more very slender roots into the pores of the ice and through the whole length of the lump. The young plant must have thrown out a considerable quantity of heat; for though the ice was, as I have said, otherwise solid, the pores through which the roots passed were enlarged to perhaps double the diameter of the fibres, but still not so much as to prevent the retention of water in them by capillary attraction.]

It does not appear that observations have been made on the special point of the development of heat in forest trees during florification, or at any other period of intense vital action; and hence an important element in the argument remains undetermined. The "circulation of the sap" commences at a very early period in the spring, and the temperature of the air in contact with trees may then be sufficiently affected by heat evolved in the vital processes of vegetation, to raise the thermometric mean of wooded countries for that season, and, of course, for the year. The determination of this point is of much greater importance to vegetable physiology than the question of the winter temperature of trees, because a slight increment of heat in the trees of a forest might so affect the atmosphere in contact with them as to make possible the growing of many plants in or near the wood which could not otherwise he reared in that climate.

The evaporation of the juices of trees and other plants is doubtless their most important thermoscopic function, and as recent observations lead to the conclusion that the quantity of moisture exhaled by vegetables has been hitherto underrated, we must ascribe to this element a higher value than has been usually assigned to it as a meteorological influence.

The exhalation and evaporation of the juices of trees, by whatever process effected, take up atmospheric heat and produce a proportional refrigeration. This effect is not less real, though to common observation less sensible, in the forest than in meadow or pasture land, and it cannot be doubted that the local temperature is considerably affected by it. But the evaporation that cools the air diffuses through it, at the same time, a medium which powerfully resists the escape of heat from the earth by radiation. Visible vapors, fogs and clouds, it is well known, prevent frosts by obstructing radiation, or rather by reflecting back again the heat radiated by the earth, just as any mechanical screen would do. On the other hand, fogs and clouds intercept the rays of the sun also, and hinder its heat from reaching the earth. The invisible vapors given out by leaves impede the passage of heat reflected and radiated by the earth and by all terrestrial objects, bat oppose much less resistance to the transmission of direct solar heat, and indeed the beams of the sun seem more scorching when received through clear air charged with uncondensed moisture than after passing through a dry atmosphere. Hence the reduction of temperature by the evaporation of moisture from vegetation, though sensible, is less than it would be if water in the gaseous state were as impervious to heat given out by the sun as to that emitted by terrestrial objects.

Total Influence of the Forest on Temperature.

It has not yet been found practicable to measure, sum up, and equate the total influence of the forest, its processes and its products, dead and living, upon temperature, and investigators differ much in their conclusions on this subject. It seems probable that in every particular case the result is, if not determined, at least so much modified by local conditions which are infinitely varied, that no general formula is applicable to the question. In the report to which I referred on page 163, Gay-Lussac says; "In my opinion we have not yet any positive proof that the forest has, in itself, any real influence on the climate of a great country, or of a particular locality. By closely examining the effects of clearing off the woods, we should perhaps find that, far from being an evil, it is an advantage; but these questions are so complicated when they are examined in a climatological point of view, that the solution of them is very difficult, not to say impossible." Becquerel, on the other hand, considers it certain that in tropical climates the destruction of the forests is accompanied with an elevation of the mean temperature, and he thinks it highly probable that it has the same effect in the temperate zones. The following is the substance of his remarks on this subject: "Forests act as frigorific causes in three ways: