Bark is the outer part of the cortex that has perished, having been cut off from nourishment by the thin hard plates of the bark-scales. In the late and early bark-formation is found a general but by no means an exact distinction between Soft and Hard Pines. In the Soft Pines the cortex remains alive for many years, adjusting itself by growth to the increasing thickness of the wood. The trunks of young trees remain smooth and without rifts. In the Hard Pines the bark-formation begins early and the trunks of young trees are covered with a scaly or rifted bark. The smooth upper trunk of older trees is invariable in Soft Pines, but in Hard Pines there are several exceptions to early bark-formation. These exceptions are easily recognized in the field, and the character is of decisive specific importance (glabra, halepensis, etc.).
Among species with early bark-formation are two forms of bark: 1, cumulative, sufficiently persistent to acquire thickness and the familiar dark gray and fuscous-brown shades of bark long exposed to the weather; 2, deciduous, constantly falling away in thin scales and exposing fresh red inner surfaces. The latter are commonly known as Red Pines, as distinguished from Black Pines with dark cumulative bark. Deciduous bark changes after some years to cumulative bark, and the upper trunk only of mature trees is red. Red Pines, although usually recognizable by their bark, are by no means constant in this character. Oecological or pathological influences may check the fall of the bark-scales, and then the distinction between the upper and lower parts of the trunk becomes lost.
SUMMARY
The various characters that have been considered in the previous pages may be classified under different heads, some of them applicable to the whole genus, others to larger or smaller groups of species.
GENERIC CHARACTERS
Several characters, quite distinct from those of other genera, are common to all the species.
- The primary leaf—appearing as a scale or bract throughout the life of the tree.
- The bud—its constant position at the nodes.
- The internode—its three distinct divisions.
- The secondary leaves—in cylindrical fascicles with a basal sheath.
- The pistillate flower—its constant nodal position and its verticillate clusters.
- The staminate flower—its constant basal position on the internode and its compact clusters.
- The cone—its clearly defined annual growths.