Skull of Domestic Ox, from a specimen in the Royal College of Surgeons.
The mouth of animals of the Ox Tribe contains, when full, thirty-two teeth. Six molars in each jaw, above, below, and on either side; and eight incisors in the lower jaw. In the upper jaw there are no incisors; but instead thereof a fibrous and elastic pad, or cushion, which covers the convex extremity of the anterior maxillary bone, and which is well worthy of observation.
The final cause of this pad (which stands in the place of upper incisor teeth) and the part it plays in the procuring of food, is thus described by Youatt. "The grass is collected and rolled together by means of the long and moveable tongue; it is firmly held between the lower cutting teeth and the pad, the cartilaginous upper lip assisting in this; and then by a sudden nodding motion of the head, the little roll of herbage is either torn or cut off, or partly both torn and cut.
"The intention of this singular method of gathering the food, it is somewhat difficult satisfactorily to explain. It is peculiar to ruminants, who have one large stomach, in which the food is kept as a kind of reservoir until it is ready for the action of the other stomachs. While it is kept there it is in a state of maceration; it is exposed to the united influence of moisture and warmth, and the consequence of this is, that a species of decomposition sometimes commences, and a vast deal of gas is extricated.
"That this should not take place in the natural process of retention and maceration, nature possibly established this mechanism for the first gathering of the food. It is impossible that half of that which is thus procured can be fairly cut through; part will be torn, and no little portion will be torn up by the roots. If cattle are observed while they are grazing, it will be seen that many a root mingles with the blades of grass; and these roots have sometimes no inconsiderable quantity of earth about them. The beast, however, seems not to regard this; he eats on, dirt and all, until his paunch is filled.
"It was designed that this earth should be gathered and swallowed; it was the meaning of this mechanism. A portion of absorbent earth is found in every soil, sufficient not only to prevent the evil that would result from occasional decomposition, by neutralizing the acid principle as rapidly as it is evolved; but, perhaps, by its presence, preventing that decomposition from taking place. Hence the eagerness with which stall-fed cattle, who have not the opportunity of plucking up the roots of grass, evince for mould. It is seldom that a cow will pass a newly-raised mole hill without nuzzling into it, and devouring a considerable portion of it. This is particularly the case where there is any degree of indigestion."
The general disposition of animals of this class, when unmolested, is inoffensive and retiring; but when excited and irritated, they are fierce and courageous, and extremely dangerous to encounter. It is a remarkable circumstance in their history, that they are generally provoked to attack at the sight of red, or any very bright and glaring colour.
Skeleton of Domestic Ox, from a specimen in the Royal College of Surgeons.