Temperature, Pressure, Pain.—The feeling of temperature, pressure, and pain is determined by different end organs in the skin. Two kinds of nerve fibers exist in the skin, which give distinct sensations of heat and cold. These nerve endings can be located by careful experimentation. There are also areas of nerve endings which are sensitive to pressure, and still others, most numerous of all, sensitive to pain.

A, isolated taste bud, from whose upper free end project the ends of the taste cells; B, supporting or protecting cell; C, sensory cell.

Taste Organs.—The surface of the tongue is folded into a number of little projections known as papillæ. These may be more easily found on your own tongue if a drop of vinegar is placed on its broad surface. In the folds, between these projections on the top and back part of the tongue, are located the organs of taste. These organs are called taste buds.

Each taste bud consists of a collection of spindle-shaped neurones, each cell tipped at its outer end with a hairlike projection. These cells send inward fibers to other cells, the fibers from which ultimately reach the brain. The sensory cells are surrounded by a number of projecting cells which are arranged in layers about them. Thus the organ in longitudinal section looks somewhat like an onion cut lengthwise.

How we Taste.—Four kinds of substances may be distinguished by the sense of taste. These are sweet, sour, bitter, and salt. Certain taste cells located near the back of the tongue are stimulated only by a bitter taste. Sweet substances are perceived by cells near the tip of the tongue, sour substances along the sides, and salt about equally all over the surface. A substance must be dissolved in fluid in order to be tasted. Many things which we believe we taste are in reality perceived by the sense of smell. Such are spicy sauces and flavors of meats and vegetables. This may easily be proved by holding the nose and chewing, with closed eyes, several different substances, such as an apple, an onion, and a raw potato.

Smell.—The sense of smell is located in the membrane lining the upper part of the nose. Here are found a large number of rod-shaped cells which are connected with the brain by means of the olfactory nerve. In order to perceive odors, it is necessary to have them diffused in the air; hence we sniff so as to draw in more air over the olfactory cells.

The Organ of Hearing.—The organ of hearing is the ear. The outer ear consists of a funnel-like organ composed largely of cartilage which is of use in collecting sound waves. This part of the ear incloses the auditory canal, which is closed at the inner end by a tightly stretched membrane, the tympanic membrane or ear drum. The function of the tympanic membrane is to receive sound waves, for all sound is caused by vibrations in the air, these vibrations being transmitted, by the means of a complicated apparatus found in the middle ear, to the real organ of hearing located in the inner ear.

Section of ear: E.M., auditory canal; Ty.M., tympanic membrane; Eu., Eustachian tube; Ty, middle ear; Coc., A.S.C., E.S.C., etc., internal ear.