On the Sponges in Chalk and Flint.—From the durability of the tissue of the flexible sponges, and the imperishable nature of those which have a siliceous or calcareous endo-skeleton or framework, their fossil remains generally occur in a fine state of preservation, and in immense quantities, in the sediments that were deposited in those parts of the ancient sea-bottoms, originally inhabited by these zoophytes. Even the relics of the keratose species, the Halichondria, whose structure consists of siliceous spines imbedded in a cartilaginous mass, are equally abundant. Sponge-spicula are everywhere met with in the chalk, flint, and greensand, and many layers in the cretaceous strata are almost entirely composed of them.
Sponges so commonly form the nuclei of the nodular flints, that some naturalists have ascribed the formation of the layers and nodules of silex in the cretaceous rocks to these zoophytes: a supposition altogether groundless,[198] The various states of mineralization in which sponges occur in the chalk give rise to many beautiful and highly instructive fossils, as we shall point out in the course of this notice. In general the zoophyte is simply invested by the flint, the pores and tubes being filled with silex, the original tissue appearing as a brown reticulated calcareous mass. In other examples the sponge has been permeated by the liquid flint, and subsequently perished; and in this manner have been formed those hollow nodules which on being broken are found to contain only a powder, consisting of siliceous spicules and fragments of silicified sponge. But in numerous instances the substance of the zoophyte is completely silicified, and the intimate structure of the original exquisitely preserved; such are many of the flint-pebbles, and moss-agates, that are mounted as brooches and other ornaments.
[198] See Wonders of Geology, p. 300. This question is fully considered in a Memoir entitled Notes of a Microscopical Examination of the Chalk and Flint of the South-East of England, &c. by the Author, in 1845.
Spongites.[199]—This name I would apply generically to those fossils which appear to be identical in structure with the ordinary marine sponges that consist of a fibro-reticulated porous mass, destitute of regular tubes or canals: the form exceedingly various.
[199] Achilleum of Schweigger.
The fossil sponges of the chalk may be divided into two groups; the cyathiforms, or cup-shaped, and the ramose, or branched. Flints inclosing the first kind, generally exhibit externally the form of the original; those containing the branched species are of irregular shapes, and except by an experienced observer, the nature of the enclosed body would not be suspected. On breaking them, the sponge is often well displayed, as in the specimen figured in [Lign. 69, fig. 2]: the surface of this fossil was covered with a white gritty powder, made up of minute needle-shaped siliceous spicula.
SPONGITES IN FLINTS.
Spongites Ramosus.—A branched sponge, sometimes from twelve to fifteen inches long, is not uncommon in the flints of the Lewes and Brighton chalk; the stems and branches are cylindrical, and the terminations of the latter are rounded and full of large pores. When completely silicified the structure can only be detected by fracture, but occasionally the sponge appears to have been saturated with liquid chalk before it was enveloped in the flint; and as it is coated with calcareous matter, it may be detached from the nodule entire.[200]
[200] In this manner I obtained the beautiful specimen (now in the British Museum) figured in my Foss. South Downs, tab, xv. fig. 11. A branch of this species is represented Pict. Atlas, pl. xxxix. fig. 12.
Spongites lobatus (sp. Fleming) is figured Pict. Atlas, pl. xxxix. fig. 6.