From this general account of the vegetable structures that may be expected to occur in the mineral kingdom, the student will in some measure be prepared for the investigation of fossil trees and plants; but for the guidance of those who are wholly unacquainted with the principles on which the Natural System of Botany adopted in this work, is founded, I am induced to present the following concise view of the principal divisions of the vegetable kingdom, though it involves some repetition.
BOTANICAL PRINCIPLES.
The following summary is given nearly in Dr. Lindley's own language:—
Botanical Principles.—One of the first things that strikes an inquirer into the structure of plants, is the fact, that while all species are capable of propagating their race, the mode in which this function is effected is essentially different in different cases. In most tribes of plants, flowers are produced, and these are succeeded by fruit, containing seed, which is shed, or scattered abroad, and grows into new individuals. But in certain families (the Cryptogamia), as Ferns, Mosses, Mushrooms, and the like, neither flowers, nor seeds properly so called, have been detected; but propagation is effected by the dispersion of grains or spores, which are usually generated in the substance of the plant, and seem to have but little analogy with true seeds. Hence the vegetable kingdom is separated into two distinct groups, namely, the flowering (Phanerogamia), and the flowerless (Cryptogamia or Agamia). As the former usually possess a highly developed system of spiral and other vessels, while the latter are either altogether destitute of them, or have them only in a few of the highest orders, and those in a peculiar state, the flowering plants are termed Vasculares, and the flowerless Cellulares. And as all the flowering, or vascular plants, when they form stems, increase by an extension of their ends, and a distension or enlargement of their circumference, but the flowerless or cellular plants form their stems simply by the addition of new matter to their points, the latter are called Acrogens, signifying increase from the summit.
Flowering plants are also for the most part furnished with respiratory or breathing organs (stomata), of which the flowerless vegetables are to a great extent destitute.
The flowering or vascular plants are also divisible into two well marked groups, namely, the Exogens, or Dicotyledons, and the Endogens, or Monocotyledons.
The Exogens (growing from without), increase by the addition of new woody matter to the outside of the stems beneath the bark; and they are further characterized by the embryo having two or more cotyledons, or seed-lobes, hence they are also called Dicotyledons; such as the Elm, Beech, &c.
The Endogens, as we have previously stated, increase by the addition of ligneous matter to the inside of their stems near the centre; and as the embryo in this class has but one cotyledon, they are likewise termed monocotyledons, as the Cane, Palm, &c. Again, exogenous plants have the young external wood connected with a central pith, by medullary processes; while endogens do not possess such a structure, having no central pith. In exogens the veins (venation) of the leaves, are disposed in meshes, like net-work, but in endogens the veins run parallel to each other.
The number of parts in the flower of an exogenous plant is usually five, or its multiples: in the endogens it is commonly three, or its multiples. In the germination, the young root of exogens is a mere extension of the radicle; but in endogens it is protruded from within.
Thus, in the flowering or vascular plants, we have two groups distinct from each other in their germination, the structure of their stems and leaves, their mode of growth, the arrangement of the parts of the flower, and in the structure of the embryo.