FOSSIL FERNS.
Filicites, or Ferns.
We now arrive at the consideration of one of the most interesting families of the vascular cryptogamia that adorned the Flora of the ancient world, and the living species of which impart beauty and elegance to the scenery of the countries where they prevail. The most essential character of these vegetables, is that of developing their fructification on the leaves; a fact familiar to every one who has even but cursorily examined the Polypody growing on our walls, or the Brake of our hedge-rows and commons. The largest species of British ferns scarcely exceed four or five feet in height; but the arborescent or tree-ferns, of warm climates, attain an altitude of from thirty to forty feet. There is too this peculiarity in the arborescent forms, that while in our indigenous species the leaves surround the stem, and incline towards the upper part of the plant, the foliage of the former bends downwards, and spreads out from the crown, or summit, into an elegant canopy.
Lign. 16. Pecopteris Sillimani; nat.
Coal Shale. Ohio.
| a. | The Stem. |
| b. | Leaf-stalk, or petiole. |
| c. | Leaf, or frond, which is bipinnate. |
| d. e. | Leaflets, or pinnæ; the upper, d, are entire; the lower, e, are pinnatifid. |
| f. | The pinnules, lobes, or segments. |
| g. | The midrib, or median vein. |
| h. | The veins. The veins are introduced in the leaflets, d; but in the lower ones, e, the midribs only are marked. |
The leaves of our branched ferns are persistent, and when shed, the markings left by their attachment to the stalk are soon obliterated. In the arborescent ferns, on the contrary, the petioles become detached from their bases, and fall entire, leaving scars or cicatrices on the stem; and these impressions are so regularly and symmetrically disposed, as to afford characters by which the trunks may be distinguished from those of other trees. The stems of the tree-ferns are therefore easily recognized in a fossil state externally, by their cylindrical forms without ramification, and by the regular disposition and peculiar character of the scars left by the separation of the petioles; and, internally by that peculiar zone, formed of bundles of ligneous tissue inclosed in sheaths, which encircles the central axis, as shown in the transverse sections in [Lign. 2], ante, [p. 62]. The leaves may be identified by the form of their segments, which are disposed with remarkable regularity, and have a peculiar mode of subdivision; and above all, by the delicacy, evenness, and distribution of the veins. There are upwards of two thousand species of living ferns, and in the fossil kingdom the number is considerable; more than two hundred have been collected from the carboniferous formation. The recent tree-ferns are confined almost exclusively to the equinoctial regions; humidity and heat being the conditions most favourable to their development (Vég. Foss. p. 141. Bd. p. 461. Wond. p. 727).
From the elegance and diversity of form of their foliage, fossil ferns are the most remarkable and attractive vegetable remains in the ancient strata. The greater number are from the coal deposits, the fern-leaves generally occurring in the schists or shales that form the roof of the beds of coal. Many of the strata are made up of carbonized fern-leaves and stems closely pressed together. The roof of a coal-mine, when newly exposed, often presents a most interesting appearance, from the abundance and variety of leaves, branches, and stems, that occur either in relief, or impressed on the dark glossy surface. The specimens selected for illustration exhibit the principal modes of venation on which the genera are founded.