This genus was instituted by M. Brongniart, for the reception of some very large fern-leaves from the shale of Hoer, in Scania, which resemble in structure the foliage of the recent Polypodium quercifolium, a native of the East Indies, and the Moluccas. One leaf was four feet wide, and the leaflets, though imperfect, were eighteen inches long.[88]
[88] Hoer is a little village, situated nearly in the centre of Scania, a province in the southern extremity of Sweden. The Chalk formation appears in several parts of this district, and Carboniferous strata at Hoeganes. To the west of Hoer, there is a range of hills, composed of ferruginous grits, micaceous sandstones, clays, and beds of quartzose conglomerate. It is in these strata that the ferns and other terrestrial plants occur, and no animal remains whatever have been found in them; their geological position appears to be between the Chalk and the Coal, but on this point nothing positive is known. The general analogy of the plants with the group forming the Flora of the Wealden, led M. Brongniart to suppose that the deposits in question belong to that formation; and M. Nillson, of Lund, who examined my collection at Brighton, recognized, among some undescribed plants from Tilgate Forest, forms that he had collected from Hoer. See "Observations sur les Végétaux Fossiles renfermés dans les Grès de Hoer en Scanie: par M. Ad. Brongniart." Ann. Sc. Nat. 1825.
Many other genera of fossil ferns have been established from the form and venation of the leaves, and are described in Brit. Foss. Flor., and other British and foreign works.
Stems of arborescent Ferns.—Notwithstanding the profusion with which the foliage of many kinds of ferns is distributed throughout the coal formation, the undoubted stems of plants of this family are rarely met with; for the numerous tribe called Sigillariæ is now removed altogether from this class. It may, however, admit of question whether much of the foliage which, from the analogy of structure, has been referred to ferns, may not have belonged to those trees; for as in the animal kingdom, so in the vegetable, distinct types of living organisms are often found blended in the lost races; and as the stems of recent tree-ferns are even more durable than their leaves, it seems impossible to account for their absence in strata, that inclose entire layers of the foliage matted together. A few fossils, supposed to possess the essential characters of recent fern-stems, have been discovered, and arranged under the following genus.
FOSSIL FERN-STEMS.
Caulopteris (fern-stem). [Lign. 31.]—Stems not channelled, marked with discoidal, oblong, or ovate scars, arranged longitudinally; vascular cicatrices numerous.
Lign. 31. Caulopteris macrodiscus. Coal.