Of course, therefore, all the ordinary unscientific conceptions of how plants feed are absolutely erroneous. Vegetable physiology, indeed, got beyond these conceptions a good hundred years ago. But it usually takes a hundred years for the world at large to make up its leeway. Trees don't suck up their nutriment by the roots, they don't derive their food from the soil, they don't need to be fed, like babies through a tube, with terrestrial solids. The solitary instance of an orchid hung up by a string in a conservatory on a piece of bark, ought to be sufficient at once to dispel for ever this strange illusion—if people ever thought; but of course they don't think—I mean Other People. The true mouths and stomachs of plants are not to be found in the roots, but in the green leaves; their true food is not sucked up from the soil, but is inhaled through tiny channels from the air; the mass of their material is carbon, as we can all see visibly to the naked eye when a log of wood is reduced to charcoal: and that carbon the leaves themselves drink in, by a thousand small green mouths, from the atmosphere around them.
But how about the juice, the sap, the qualities of the soil, the manure required? is the incredulous cry of Other People. What is the use of the roots, and especially of the rootlets, if they are not the mouths and supply-tubes of the plants? Well, I plainly perceive I can get 'no forrarder,' like the farmer with his claret, till I've answered that question, provisionally at least; so I will say here at once, without further ado—the plant requires drink as well as food, and the roots are the mouths that supply it with water. They also suck up a few other things as well, which are necessary indeed, but far from forming the bulk of the nutriment. Many plants, however, don't need any roots at all, while none can get on without leaves as mouths and stomachs. That is to say, no true plantlike plants, for some parasitic plants are practically, to all intents and purposes, animals. To put it briefly, every plant has one set of aerial mouths to suck in carbon, and many plants have another set of subterranean mouths as well, to suck up water and mineral constituents.
Have you ever grown mustard and cress in the window on a piece of flannel? If so, that's a capital practical example of the comparative unimportance of soil, except as a means of supplying moisture. You put your flannel in a soup-plate by the dining-room window; you keep it well wet, and you lay the seeds of the cress on top of it. The young plants, being supplied with water by their roots, and with carbon by the air around, have all the little they need below, and grow and thrive in these conditions wonderfully. But if you were to cover them up with an air-tight glass case, so as to exclude fresh air, they'd shrivel up at once for want of carbon, which is their solid food, as water is their liquid.
The way the plant really eats is little known to gardeners, but very interesting. All over the lower surface of the green leaf lie scattered dozens of tiny mouths or apertures, each of them guarded by two small pursed-up lips which have a ridiculously human appearance when seen through a simple microscope. When the conditions of air and moisture are favourable, these lips open visible to admit gases; and then the tiny mouths suck in carbonic acid in abundance from the air around then. A series of pipes conveys the gaseous food thus supplied to the upper surface of the leaf, where the sunlight falls full upon it. Now, the cells of the leaf contain a peculiar green digestive material, which I regret to say has no simpler or more cheerful name than chlorophyll; and where the sunlight plays upon this mysterious chlorophyll, it severs the oxygen from the carbon in the carbonic acid, turns the free gas loose upon the atmosphere once more through the tiny mouths, and retains the severed carbon intact in its own tissues. That is the whole process of feeding in plants: they eat carbonic acid, digest it in their leaves, get rid of the oxygen with which it was formerly combined, and keep the carbon stored up for their own purposes.
Life as a whole depends entirely upon this property of chlorophyll; for every atom of organic matter in your body or mine was originally so manufactured by sunlight in the leaves of some plant from which, directly or indirectly, we derive it.
To be sure, in order to make up the various substances which compose their tissues—to build up their wood, their leaves, their fruits, their blossoms—plants require hydrogen, nitrogen, and even small quantities of oxygen as well; but these various materials are sufficiently supplied in the water which is taken up by the roots, and they really contribute very little indeed to the bulk of the tree, which consists for the most part of almost pure carbon. If you were to take a thoroughly dry piece of wood, and then drive off from it by heat these extraneous matters, you would find that the remainder, the pure charcoal, formed the bulk of the weight, the rest being for the most part very light and gaseous. Briefly put, plants are mostly carbon and water, and the carbon which forms their solid part is extracted direct from the air around them.
How does it come about then that a careless world in general, and more especially the happy-go-lucky race of gardeners and farmers in particular, who have to deal so much with plants in their practical aspect, always attach so great importance to root, soil, manure, minerals, and so little to the real gaseous food stuff of which their crops are, in fact, composed? Why does Hodge, who is so strong on grain and guano, know absolutely nothing about carbonic acid? That seems at first sight a difficult question to meet. But I think we can meet it with a simple analogy.
Oxygen is an absolute necessary of human life. Even food itself is hardly so important an element in our daily existence; for Succi, Dr. Tanner, the prophet Elijah, and other adventurous souls too numerous to mention, have abundantly shown us that a man can do without food altogether for forty days at a stretch, while he can't do without oxygen for a single minute. Cut off his supply of that life-supporting gas, choke him, or suffocate him, or place him in an atmosphere of pure carbonic acid, or hold his head in a bucket of water, and he dies at once. Yet, except in mines or submarine tunnels, nobody ever takes into account practically this most important factor in human and animal life. We toil for bread, but we ignore the supply of oxygen. And why? Simply because oxygen is universally diffused everywhere. It costs nothing. Only in the Black Hole of Calcutta or in a broken tunnel shaft do men ever begin to find themselves practically short of that life-sustaining gas, and then they know the want of it far sooner and far more sharply than they know the want of food on a shipwreck raft, or the want of water in the thirsty desert. Yet antiquity never even heard of oxygen. A prime necessary of life passed unnoticed for ages in human history, only because there was abundance of it to be had everywhere.
Now it isn't quite the same, I admit, with the carbonaceous food of plants. Carbonic acid isn't quite so universally distributed as oxygen, nor can every plant always get as much as it wants of it. I shall show by-and-by that a real struggle for food takes place between plants, exactly as it takes place between animals; and that certain plants, like Oliver Twist in the workhouse, never practically get enough to eat. Still, carbonic acid is present in very large quantities in the air in most situations, and is freely brought by the wind to all the open spaces which alone man uses for his crops and his gardening. The most important element in the food of plants is thus in effect almost everywhere available, especially from the point of view of the mere practical everyday human agriculturist. The wind that bloweth where it listeth brings fresh supplies of carbon on its wings with every breeze to the mouths and throats of the greedy and eager plants that long to absorb it.
It is quite otherwise, however, with the soil and its constituents. Land, we all know—or if we don't, it isn't the fault of Mr. George and Mr. A.R. Wallace—land is 'naturally limited in quantity.' Every plant therefore struggles for a foothold in the soil far more fiercely and far more tenaciously than it struggles for its share in the free air of heaven. Your plant is a land-grabber of Rob Roy proclivities; it believes in a fair fight and no favour. A sufficient supply of food it almost takes for granted, if only it can once gain a sufficient ground-space. But other plants are competing with it, tooth and nail (if plants may be permitted by courtesy those metaphorical adjuncts), for their share of the soil, like crofters or socialists; every spare inch of earth is permeated and pervaded with matted fibres; and each is striving to withdraw from each the small modicum of moisture, mineral matter, and manure for which all alike are eagerly battling.