Diagram 7.—Showing how the Digestive Canal is Lengthened.
Diagram 8.—Cross-section of the Digestive Tube.
The digestive canal has, however, another function. The cells which compose it have not only to secrete juices, to convert the food into a usable form; they have then to absorb it. The nearer a particle of food is to the wall of cells, the sooner it is reached by these juices, and the less chance there is of useful material being swept away and lost. In view of this fact, along certain tracts the digestive canal is folded inwards, and there are projections, which increase the number of cells to secrete and their opportunities of absorption. ([See Diagram 8.])
Diagram 9.—Showing how Glands arise.
Here again we have an illustration of a constantly recurring need, with a device for meeting it—increase of surface without increase of bulk. We met with it before in the cellular system; we shall meet with it again in glands, lungs, and brain, at least. The importance of a device for gaining this end is apparent when one remembers what the comparative value of surface and bulk is to an animal, and that, while surface increases by the square, bulk increases by the cube.
The principle is pressed to an extreme, together with the allied principle of division of labour, in glands. The object of these is to increase the number of secreting cells, and, as they are delicate, to keep them protected from contact with coarse particles of food. And, in order that nothing may interfere with their efficiency, they are absolved from the duty of absorbing. Hence tubes grow out from the cavity of the alimentary canal lined with the same cells, but, as no food ever enters, the cells which line them devote themselves entirely to pouring out digestive juices. Glands differ considerably in structure and in the liquids which they secrete. Some are very small; some, like the liver, very large. In some the tube is very short, in some long, coiled and branched, and sometimes the gland is connected with the surface by more or less of a duct. Some glands only secrete one enzyme, some several. In each, however, the principle is that shown in [Diagram 9], no matter how its structure is masked by the bloodvessels and supporting cells or connective tissue which envelop it.
After a meal, or, rather, when the process of digestion is over and the animal is beginning to think about its next, the gland cells start preparing their enzyme. There is great activity in the nucleus, and granules stream out from it towards the lumen of the gland in much the same way, to take a homely illustration, as bubbles in some effervescing drink form at the bottom of the tumbler and rise till the surface is covered with foam. At the right moment these granules are discharged, just as the bubbles on the surface of a liquid break at a slight jog. They are usually not the ferment or enzyme, but its precursor, a substance which only turns into the ferment when it gets outside the cells. The ferments, when formed, are very peculiar substances about which we should like the chemist to tell us more, though great advances have been made in our knowledge of them lately.
Among other peculiarities, one may mention that, though they will keep indefinitely if bottled, they are easily destroyed by too extreme a temperature or too acid or alkaline surroundings, that their composition is entirely unknown, and, strangest of all, that they do not become used up. A given amount of rennet will clot any amount of milk within reasonable limits, and yet remain rennet. The clergyman has been quoted as an illustration of the action of a ferment, and he makes a good one. He can make any number of suitable men and women into married couples, and yet his own state is unchanged.