The outer and hinder border of the nostril is made by another bone named the maxillary bone, which is usually much shorter than the premaxillary. It contains the hindermost teeth, which rarely differ from those in front, except in sometimes being smaller.
The nasal bones, which always make the upper and hinder border of the nostrils, meet each other above them, in the middle line of the beak.
Showing that the extremity of the jaws in Rhamphorhynchus was sheathed in horn as in the giant Kingfisher, since the jaws similarly gape in front.
The hyoid bones are below the lower jaw in the Pterodactyle.
The nostrils are unusually large in the Lias genus named Dimorphodon, and small in species of the genus Rhamphorhynchus from Solenhofen. Such differences result from the relative dimensions and proportions of these three bones which margin the nasal vacuity, and by varying growth of their front margins or of their hinder margins govern the form of the snout.
The jaws are most massive in the genera known from the Wealden beds to the Chalk. The palatal surface is commonly flat or convex, and often marked by an elevated median ridge which corresponds to a groove in the lower jaw, though the median ridge sometimes divides the palate into two parallel concave channels. The jaw is margined with teeth which are rarely fewer than ten or more than twenty on each side. They are sharp, compressed from side to side, curved inward, and never have a saw-like edge on the back and front margins. No teeth occur upon the bones of the palate.
In most birds there is a large vacuity in the side of the head between the nostril and the orbit of the eye, partly separated from it by the bone which carries the duct for tears named the lachrymal bone. The same preorbital vacuity is present in all long-tailed Pterodactyles, though it is either less completely defined or absent in the group with short tails. It affords excellent distinctive characters for defining the genera. In the long-tailed genus Scaphognathus from Solenhofen this preorbital opening is much larger than the nostril, while in Dimorphodon these vacuities are of about equal size. Rhamphorhynchus is distinguished by the small size of the antorbital vacuity, which is placed lower than the nostril on the side of the face. The aperture is always imperfectly defined in Pterodactylus, and is a relatively small vacuity compared with the long nostril. In Ptenodracon the antorbital vacuity appears to have no existence separate from the nostril which adjoins the eye hole. And so far as is known at present there is no lateral opening in advance of the eye in the skull in any Ornithosaur from Cretaceous rocks, though the toothless Ornithostoma is the only genus with the skull complete. When a separate antorbital vacuity exists, it is bordered by the maxillary bone in front, and by the malar bone behind. The prefrontal bone is at its upper angle. That bone is known in a separate state in reptiles and, I think, in monotreme mammals. Its identity is soon lost in the mammal, and its function in the skull is different from the corresponding bone in Pterodactyles.