Section 131. A cross section of the spinal cord is shown in Figure 6, [Sheet 8]. It is a cylinder, almost bisected by a dorsal (d.f.) and a ventral (v.f.) fissure. Through its centre runs a central canal (c.c.), continuous with the brain ventricles, and lined by ciliated epithelium. The spinal cord consists of an outer portion, mainly of nervous fibres, the white matter, and of inner, ganglionated, and more highly vascular grey matter. (In the cerebrum the grey matter is external, and the white internal.) The cord, like the brain, is surrounded by a vascular fibrous investment, and protected from concussion by a serous fluid. The nerves which emerge from the vertebral column between the vertebrae, arise, unlike the cranial nerves, by two roots. The dorsal of these, the sensory root (d.n.), has a swelling upon it, the dorsal ganglion, and-- by experiments upon living animals-- has been shown to contain only afferent fibres; the ventral, the motor root, is without a ganglion, and entirely or mainly motor. The two unite outside the cord, and thereafter the spinal nerves are both sensory and motor.
Section 132. Besides the great mass of brain and spinal cord (cerebro-spinal axis), there is, on either side of the dorsal wall of the body cavity, a sympathetic nervous chain. The nerve fibres of this system, like the nerve fibres of invertebrates, are non-medullated. It may be seen as a greyish thread running close by the common carotid in the neck (sym., [Figure 1]); it then runs over the heads of the ribs in the thorax and close beside the dorsal aorta in the abdominal region. In the anterior region of the neck it dilates to form a superior cervical ganglion, and opposite the first rib it forms an inferior cervical ganglion. Thence, backwards, there is a ganglion on each sympathetic chain opposite each spinal nerve, and the two exchange fibres through a thread, the ramus communicans. To the sympathetic chain is delegated much of the routine work of reflex control of the bloodvessels and other viscera, which would otherwise fall upon the spinal cord.
Section 133. There are eight cervical (spinal) nerves, one in front of the atlas, and one behind each of the cervical vertebrae. The last four and the first thoracic (spinal) contribute to a leash of nerves running out to the fore limb, the brachial plexus (plexus, literally network, but here meaning a plaited cord). The fourth cervical also sends down a phrenic nerve (p.n., [Figure 1]), along by the external jugular vein and the superior caval vein to the diaphragm. The last three lumbar and the sacral nerves form a sacral plexus, supplying the hind limb.
Section 134. From the sympathetic in the hinder region of the thorax a nerve, the great splanchnic nerve, arises, and runs, back to a ganglionated nervous network, just behind the coeliac artery, into which the vagus also enters; this is the coeliac ganglion, and together with a similar superior mesenteric ganglion around the corresponding artery, makes up a subsidiary visceral nervous network, the solar plexus. A similar and smaller nervous tangle, bearing an inferior mesenteric ganglion, lies near the inferior mesenteric artery.
Section 135. Finally, we may note the pineal gland and the pituitary body, as remarkable appendages above and below the thalamencephalon. Their function, if they have a function, is altogether unknown. Probably, they are inherited from ancestors to whom they were of value. Such structures are called reduced or vestigial structures, and among other instances are the clavicles of the rabbit, the hair on human limbs, the little pulpy nodule in the corner of the human eye, representing the rabbit's third eyelid, and the caudal vertebrae at the end of the human spinal column. In certain lowly reptiles, in the lampreys, and especially in a peculiar New Zealand lizard, the pineal gland has the most convincing resemblance to an eye, both in its general build and in the microscopic structure of its elements; and it seems now more than probable that this little vascular pimple in our brains is a relic of a third and median eye possessed by ancestral vertebrata. The pituitary body is probably equivalent to a ciliated pit we shall describe in the lancelet (Amphioxus).
8. _Renal and Reproductive Organs_
Section 136. We have now really completed our survey of the individual animal's mechanism. But no animal that was merely complete in itself would be long sanctioned by nature. For an animal species to survive, there must evidently, also, be proper provision for the production of young, and the preservation of the species as well as of the individual. Hence in an animal's physiology and psychology we meet with a vast amount of unselfish provision, and its structure and happiness are more essentially dependent on the good of its kind than on its narrow personal advantage. The mammalia probably owe their present dominant position in the animal kingdom to the exceptional sacrifices made by them for their young. Instead of laying eggs and abandoning them before or soon after hatching, the females retain the eggs within their bodies until the development of the young is complete, and thereafter associate with them for the purposes of nourishment, protection, and education. In the matter of the tail, for instance, already noted, the individual rabbit incurs the disadvantage of conspicuousness for the rear, in order to further the safety of the young.
Section 137. The female organs of reproduction are shown in [Sheet 10]. The essential organ is the ovary (ov.), in which the ova (eggs) are formed. [Figure 3] gives an enlarged and still more diagrammatic rendering of the ovary. There is a supporting ground mass, or stroma, into which numerous bloodvessels and nerves enter and break up. The ova appear first as small cells in the external substance of the ovary (as at 1), and move inward (2 and 3), surrounded by a number of sister cells, which afford them nourishment. At (4) an ovum with its surrounding group of cells is more distinct and near the centre of the ovary; a fluid is appearing within the ovisac as the development proceeds. (5) is a much more mature ovisac or Graafian follicle.