And it should be clear to the reader that given these elemental facts of life, given growth and death and reproduction with individual variation in a world that changes, life must change in this way, modification and differentiation must occur, old species must disappear, and new ones appear. We have chosen for our instance here a familiar sort of animal, but what is true of furry beasts in snow and ice is true of all life, and equally true of the soft jellies and simple beginnings that flowed and crawled for hundreds of millions of years between the tidal levels and in the shallow, warm waters of the Proterozoic seas.
The early life of the early world, when the blazing sun rose and set in only a quarter of the time it now takes, when the warm seas poured in great tides over the sandy and muddy shores of the rocky lands and the air was full of clouds and steam, must have been modified and varied and species must have developed at a great pace. Life was probably as swift and short as the days and years; the generations, which natural selection picked over, followed one another in rapid succession.
Natural selection is a slower process with man than with any other creature. It takes twenty years or more before an ordinary human being in western Europe grows up and reproduces. In the case of most animals the new generation is on trial in a year or less. With such simple and lowly beings, however, as first appeared in the primordial seas, growth and reproduction was probably a matter of a few brief hours or even of a few brief minutes. Modification and differentiation of species must accordingly have been extremely rapid, and life had already developed a very great variety of widely contrasted forms before it began to leave traces in the rocks. The Record of the Rocks does not begin, therefore, with any group of closely related forms from which all subsequent and existing creatures are descended. It begins in the midst of the game, with nearly every main division of the animal kingdom already represented.[9] Plants are already plants, and animals animals. The curtain rises on a drama in the sea that has already begun, and has been going on for some time. The brachiopods are discovered already in their shells, accepting and consuming much the same sort of food that oysters and mussels do now; the great water scorpions crawl among the seaweeds, the trilobites roll up into balls and unroll and scuttle away. In that ancient mud and among those early weeds there was probably as rich and abundant and active a life of infusoria and the like as one finds in a drop of ditch-water to-day. In the ocean waters, too, down to the utmost downward limit to which light could filter, then as now, there was an abundance of minute and translucent, and in many cases phosphorescent, beings.
But though the ocean and intertidal waters already swarmed with life, the land above the high-tide line was still, so far as we can guess, a stony wilderness without a trace of life.
IV
THE INVASION OF THE DRY LAND BY LIFE
§ 1. Life and Water. § 2. The Earliest Animals.
§ 1
WHEREVER the shore line ran there was life, and that life went on in and by and with water as its home, its medium, and its fundamental necessity.
The first jelly-like beginnings of life must have perished whenever they got out of the water, as jelly-fish dry up and perish on our beaches to-day. Drying up was the fatal thing for life in those days, against which at first it had no protection. But in a world of rain-pools and shallow seas and tides, any variation that enabled a living thing to hold out and keep its moisture during hours of low tide of drought met with every encouragement in the circumstances of the time. There must have been a constant risk of stranding. And, on the other hand, life had to keep rather near the shore and beaches in the shallows because it had need of air (dissolved of course in the water) and light.
No creature can breathe, no creature can digest its food, without water. We talk of breathing air, but what all living things really do is to breathe oxygen dissolved in water. The air we ourselves breathe must first be dissolved in the moisture in our lungs; and all our food must be liquefied before it can be assimilated. Water-living creatures which are always under water, wave the freely exposed gills by which they breathe in that water, and extract the air dissolved in it. But a creature that is to be exposed for any time out of the water, must have its body and its breathing apparatus protected from drying up. Before the seaweeds could creep up out of the Early Palæozoic seas into the intertidal line of the beach, they had to develop a tougher outer skin to hold their moisture. Before the ancestor of the sea scorpion could survive being left by the tide it had to develop its casing and armour. The trilobites probably developed their tough covering and rolled up into balls, far less as a protection against each other and any other enemies they may have possessed, than as a precaution against drying. And when presently, as we ascend the Palæozoic rocks, the fish appear, first of all the backboned or vertebrated animals, it is evident that a number of them are already adapted by the protection of their gills with gill covers and by a sort of primitive lung swimming-bladder, to face the same risk of temporary stranding.